Gnaphosa koponeni, Marusik, Yuri M. & Omelko, Mikhail M., 2014

Marusik, Yuri M. & Omelko, Mikhail M., 2014, A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804, Zootaxa 3894 (1) : -

Gnaphosa koponeni sp. n.

Figs 1–9 , Map 1.

Material examined. Holotype ♂ ( ISEA), Russia, SE Tuva, East of Tannu-Ola Mt. Range, 20 km of Khol’-Oozhu Vill., 50°50’N 94°19’E, mountain Larix -moss-stony forest-tundra, 2175 m, 8– 16.06.1995 (D.V. Logunov). Paratypes: 1♂ 2♀ ( ISEA), together with holotype; 4♂ ( ZMUT), Tuva, Tannu-Ola, S slope 50°50’N, 94°18’E, 2120 m, Pinus cembra-Larix forest, 8– 17.06.1995 (S. Koponen); 2♂ ( ZMUT), West Sayan, Oiskiy Pass, 1650 m, alpine tundra slope, 21.06.1995 (S. Koponen).

Etymology. The species name is a patronym honouring its collector, our friend and colleague Seppo Koponen, University of Turku, Finland.

Diagnosis. Males of the new species are most similar to G. wiehlei . The two species can be distinguished by a long tibial apophysis, large, widely spaced teeth at base of the embolus and distinct tapering of the terminal half of embolus (in prolateral view) in the new species (shorter tibial apophysis, one strong tooth (spur) on prolateral part of embolus, and gradual tapering of embolus in G. w i e h l e i). Males of the two sibling species can be separated on the basis of tibia I spination: the new species has only 2 apical spines, while G. wiehlei has 2 prolateral, 2 pairs of ventral and 2 apical spines ( Fig. 21 ). Females of the new species are most similar to G. sticta in well developed bulges of the fovea, but can be distinguished by the wider scape in the new species, which is larger than the fovea bulge (smaller than the bulge in G. s t i c t a), larger glands of receptacles bent posteriorly (shorter and horizontally spread glands in G. sticta ) and shape of lateral foveal margins: without pocket in new species and distinct pocket in G. sticta . Females of G. koponeni sp. n. differ from G. wiehlei by undeveloped bulges in G. wiehlei , short lateral foveal margins without pocket (long lateral margins with distinct pocket in G. wiehlei ). Both sexes of the new species differ from G. wiehlei by having dark-brown coloration (light yellow in G. wiehlei ).

Description. Male. Total length 6.35. Carapace 3.1 long, 2.4 wide. Carapace dark brown without pattern, fovea black. Sternum, labium, maxillae dark brown. Abdomen dark brown. Legs brown without spots. Legs spination: femora I 2 d, 1p; II 2 d, 2r; III 2 d, 2p, 2r; IV 2 d, 2p, 2r; patellae: III 1 r, other without spines; tibiae I 2 apical; II 2 apical; III 1 d, 4p, 3r, 2- 1v and 2 apical; IV 1 d, 3p, 4r, 2- 2v and 2 apical; metatarsi I 1- 1v; II 1- 1v; III 4 p, 4r, 2-0 (2-1)v and 4 apical; IV 3 p, 4r, 2- 1v and 6 apical. Length of legs joints as shown in Table 1 .

Palp as in Figs 1–6 ; tibial apophysis as long as tibia, sharply pointed, undivided; retrobasal margin of cymbium bulging (convex) (Bm); median apophysis widened; embolus long, with strong teeth (Tt) on retrolateral part and angle (Ea) in mid part, after which the embolus tapers.

Female. Total length 9.9. Carapace 3.75 long, 2.7 wide. Coloration as in males. Legs spination: femora I 2 d, 1p; II 2 d, 1p; III 2 (1)d. 2p, 2r; IV 2 d, 1p, 1r; patellae: III 1 r, other without spines; tibiae II 1 apical; III 1 d, 4p, 3r, 2- 2v and 2 apical; IV 4 p, 4r, 2- 2v and 2 apical; metatarsi I 1- 1v; II 1- 1v; III 4 p, 3r, 2- 1v and 4 apical; IV 3 p, 4r, 2- 2v and 6 apical. Length of legs joints as shown in Table 2 .

Epigyne as in Figs 7–9 ; scape large, wider than long, with concave posterior margin, pocket of scape (Ps) longer than wide; fovea with pair of large bulges (Fb) delimited by posterior furrows (Pf); lateral margins (Lm) short with almost undeveloped pocket (Mp); receptacles (Re) stretched almost horizontally; glands (Rg) relatively long, bent posteriorly.

Distribution. The new species is known from the south part of the Krasnoyarsk Province and Tuva. Its occurrence in adjacent Mongolia is very likely. Some of the female records of G. sticta from Tuva may refer to this species.

Figures

FIGURES 1 – 6. Male palp of Gnaphosa koponeni sp. n. 1 – 2 palp, ventral and retrolateral; 3 – 6 bulbus, prolateral, ventral, from above and retrolateral. Scale = 0.2 mm. Abbreviations: Bm— bulging margin of cymbium, Ea— angle of embolus, Eb— base of embolus, Tt— strong teeth of embolus.

FIGURES 7 – 15. Epigynes of Gnaphosa koponeni sp. n. (7 – 9), G. wiehlei (10 – 12) and G. sticta (13 – 15, from Finland). 7, 10, 13 intact, ventral; 8, 11, 14 macerated, ventral; 9, 12, 15 macerated, dorsal. Scale = 0.2 mm. Abbreviations: Fb— bulge of fovea, Id— insemination duct, Lm— lateral margin, Mp— pocket of lateral margin, Pf— posterior furrow, Ps— pocket of scape, Re— receptacle, Rg— gland of receptacle.

FIGURES 16 – 21. Male palp (16 – 20) and leg I (21) of Gnaphosa wiehlei. 16, 19 ventral; 17, 21 prolateral; 18, 20 retrolateral. Scale = 0.2 mm if not otherwise indicated. Abbreviations: Sp— spur of embolus.

Tables

TABLE 1. Length of leg joints in male.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
I	2.2	1.2	1.55	1.4	1.15	7.5
II	1.95	0.9	1.55	1.2	1.05	6.65
III	1.75	0.9	1.35	1.45	1.05	6.5
IV	2.2	1.2	1.85	2.25	1.25	8.75

TABLE 2. Length of leg joints in female.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
I	2.4	1.3	1.75	1.25	1.1	7.8
II	2.1	1.2	1.6	1.25	1	7.15
III	1.75	1	1.3	1.55	1.05	6.65
IV	2.55	1.1	1.85	2.35	1.3	9.15