Scirtes flavoguttatus, Champion, B, 1918

Yoshitomi, Hiroyuki & Ruta, Rafał, 2010, Revision of the Scirtes flavoguttatus species-group (Coleoptera: Scirtidae: Scirtinae) 2467, Zootaxa 2467 (1), pp. 1-74 : 72-74

Key to species of Scirtes flavoguttatus species-group

1. Apical portion of elytra with excitator (females) .......................................................................................................... 2

- Apical portion of elytra without excitator..................................................................................................................... 4

2. Elytra uniformly brownish-black, excitator oblique ..................................................................................................... 3

- Elytra dark brown to black with faint pattern consisting of spots and transverse bands ................................................ ........................................................................................................... unidentifiable females cf. Scirtes luteomaculatus

3. Pronotum uniformly brownish-black ..................................................................... Scirtes laosensis Yoshitomi & Ruta

- Sides of pronotum lighter, yellowish ........................................................................ Scirtes niisatoi Yoshitomi & Ruta

4. Elytra unicolour or gradually darkening apically ......................................................................................................... 5

- Elytra bicolour, often with contrasting, usually black and yellow, pattern ................................................................. 15

5. Elytra bright red ............................................................................................................... Scirtes rufotinctus Champion

- Elytra brown to brownish-black.................................................................................................................................... 6

6. Pronotum yellow to red, elytra contrasting dark brown............................................................................................... 7

- Pronotum and elytra of similar colour .......................................................................................................................... 9

7. Parameres longer than penis ....................................................................................... Scirtes sakaii Yoshitomi & Ruta

- Parameres not longer than penis ................................................................................................................................... 8

8. Parameres distinctly shorter than penis .................................................................... Scirtes curvans Yoshitomi & Ruta

- Parameres as long as penis ................................................................................................... Scirtes dichrous Champion

9. Elytra uniformly brown............................................................................................................................................... 10

- Elytra slightly darkened apically................................................................................................................................. 15

10. Apical portion of penis bifid, consisting of narrow struts, hidden behind parameres… Scirtes patkainus Champion

- Apical portion of penis undivided, wide, well developed........................................................................................... 11

11. Larger species, TL over 3.2 mm (usually over 3.5 mm)............................................................................................. 12

- Smaller species, TL not exceeding 3.1 mm ............................................................................................................... 14

12. Dorsum uniformly light brown, male genitalia as in Fig. 39 ............................... Scirtes brancuccii Yoshitomi & Ruta

- Dorsum uniformly dark brown ................................................................................................................................... 13

13. Parameres long, divided basally, male genitalia as in Fig. 34 , antennae subserricorn… Scirtes crassicornis Champion

- Parameres short, divided apically, male genitalia as in Fig. 33 , antennae filiform ........... Scirtes sulcigerus Champion

14. Elytra with sparser punctation, punctures separated by more than diameter of a puncture; penis much longer than parameres ( Fig. 38 ) ............................................................................................. Scirtes sabahensis Yoshitomi & Ruta

- Elytra with denser punctation, punctures separated by less than diameter of a puncture; penis not longer than parameres .................................................................................................................................................................... 15

15. Parameres regularly curved ( Fig. 37D ).................................................................. Scirtes cameroni Yoshitomi & Ruta

- Parameres straight, only apices are incurved ( Fig. 36D ) ................................................. Scirtes malayanus Champion

16. Parameres curved laterally, with three pairs of claw-like projections; caudal part of penis membranous ..................... .................................................................................................................................. Scirtes maramagensis Klausnitzer

- Parameres curved inwardly, pointed simply; caudal part of penis sclerotized, distinctly projecting ............................. .............................................................................................................................. Scirtes balehensis Yoshitomi & Ruta

17. Each elytron with a large dark spot............................................................................................................................. 18

- Each elytron with a pattern consisting of numerous spots or longitudinal stripes...................................................... 20

18. Basal part and apex of elytron brownish, central portion brownish black, not contrasting with remaining portion of elytron ............................................................................................................... Scirtes tamdaoensis Yoshitomi & Ruta

- Elytra yellow, each with a contrasting black spot ....................................................................................................... 19

19. Black spot on elytra relatively small, situated in apical portion of elytra… Scirtes nigrofasciatus Yoshitomi & Ruta

- Black area on elytra large, covering most of posterior half of elytra ............................ Scirtes satoi Yoshitomi & Ruta

20. Elytra yellow with black pattern ................................................................................................................................. 21

- Elytra black with yellow pattern ................................................................................................................................. 25

21. Elytral pattern indistinct, consisting of wide transverse dark stripes.......................................................................... 22

- Elytral pattern distinct ................................................................................................................................................. 23

22. Pronotum yellow, humeri yellow, antennomeres 4–11 dark brown ................... Scirtes malaccanus Yoshitomi & Ruta

- Pronotum dark brown, humeri with brown oval area, antennae yellow ................ Scirtes malcolmi Yoshitomi & Ruta

23. Each elytron with two large black areas, an oval one in the basal half and a slightly larger and subtriangular one in posterior half ....................................................................................................................... Scirtes warchalowskii Ruta

- Black pattern different................................................................................................................................................. 24

24. Pronotum yellow, black pattern consisting of a transverse band in the middle of each elytron and a subtriangular area in apical portion of elytra, suture yellow Scirtes kodadai Yoshitomi & Ruta

- Pronotum dark brown, black pattern consisting of basal spot, transverse band of four elongated spots in the middle of elytra, and an elongate rectangular area in apical portion of elytra ..................... Scirtes bidentatus Yoshitomi & Ruta

25. Elytra with a longitudinal yellow stripes running along the suture and lateral margins of elytra, fused at apex ....... 26

- Elytral colour pattern consisting of numerous yellow spots ....................................................................................... 27

26. Lateral stripe continuous ............... Scirtes niisatoi Yoshitomi & Ruta and Scirtes chiangmaiensis Yoshitomi & Ruta , differing in the morphology of male genitalia ( Figs. 26 , 27 )

- Lateral stripe interrupted before the middle of its length and in the posterior half ........................................................ ................................................................................................................................ Scirtes laosensis Yoshitomi & Ruta

27. Each elytron with no more than 8 yellow spots .......................................................................................................... 27

- Each elytron with more than 8 yellow spots or pattern consisting of short longitudinal stripes ................................ 31

28. Each elytron with 5 yellow spots ............................................................................................................................... 29

- Each elytron with 7–8 yellow spots ............................................................................................................................ 30

29. Larger (TL 3.43 mm), more elongate; antennae shorter and more slender; antennomeres 1–2 testaceous; elytra less convex, yellow spots on elytra larger, male unknown ................................................... Scirtes flavonotatus Champion

- Smaller (TL 2.80–2.85 mm), less elongate; antennae longer, stouter, antennomeres 1–3 testaceous; elytra more convex, yellow spots on elytra smaller ............................................................................. Scirtes decemnotatus Champion

30. Body black, 7–8 spots on each elytron, male genitalia as in Fig. 5 .............................. Scirtes flavoguttatus Champion

- Body brownish, 7 spots on each elytron, male genitalia as in Fig. 7 ........ Scirtes septemmaculatus Yoshitomi & Ruta

31. Colour patterns on each elytron consisting of yellow elytral base, two transverse rows of yellow spots – in the basal half, and in the middle of the length of elytra and larger apical spot.......................................................................... 32

- Elytral colour pattern different.................................................................................................................................... 34

32. Elytra with two transverse bands consisting of oval spots, pronotum uniformly yellow, male genitalia as in the Fig. 8 ............................................................................................................................................... Scirtes luteomaculatus Pic

- Elytra with two transverse bands consisting of elongated spots, pronotum darkened in central portion ................... 33

33. Spots on elytra in transverse bands more elongated, male genitalia as in Fig. 9 .... Scirtes teruhisai Yoshitomi & Ruta

- Spots on elytra in transverse bands less elongated, male genitalia as in Fig. 10 .......... Scirtes maxi Yoshitomi & Ruta

34. Elytral colour pattern consisting of yellow or orange basal portion, 4–5 spots arranged in transverse band in the middle of the elytra, an elongated subapical spot (absent in philippinensis ) and apical subquadrate spot....................... 35

- Elytral colour pattern different, median transverse band lacking or not consisting of spots ...................................... 38

35. Basal portion of elytra orange, elytral spots yellow.................................................................................................... 36

- Both basal portion of elytra and elytral spots yellow.................................................................................................. 37

36. Elytra with an elongated subapical spot, suture orange ........................................................... Scirtes basifasciatus Pic

- Elytra without an elongated subapical spot, suture black ............................... Scirtes philippinensis Yoshitomi & Ruta

37. Elytra pale brown, apical spot with an acute antero-lateral angle ..................... Scirtes schawalleri Yoshitomi & Ruta

- Elytra brownish-black, apical spot with a rounded antero-lateral angle ..... Scirtes schawalleroides Yoshitomi & Ruta

38. Basal 1/3 of elytra yellow with black humeral portion, apices of elytra yellow............................................................. ......................................................................................................................... Scirtes tanahratensis Yoshitomi & Ruta

- Elytral colour pattern different.................................................................................................................................... 39

39. Elytra with a longitudinal stripe in middle ............................................................................... Scirtes curtelineatus Pic

- Elytra black with yellow base, a wide transverse stripe in the middle and an oval apical spot...................................... ........................................................................................................................................ Scirtes flavocinctus Champion

Figures

FIGURE 5. Scirtes flavoguttatus Champion, holotype, male. A) sternites I–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect, F) ditto in dorsal aspect, G) ditto in lateral aspect. Abbreviations: p, penis; t, tegmen.

FIGURE 7. Scirtes septemmaculatus sp. n., paratype, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect.

FIGURE 8. Scirtes luteomaculatus Pic, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect.

FIGURE 9. Scirtes teruhisai sp. n., paratype, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect. Abbreviations: p, penis; t, tegmen.

FIGURE 10. Scirtes maxi sp. n., holotype, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect.

FIGURE 26. Scirtes chiangmaiensis sp. n., holotype, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect, F) ditto in lateral aspect.

FIGURE 27. Scirtes niisatoi sp. n., holotype, male. A) tegmen and penis in ventral aspect, B) ditto in lateral aspect. Abbreviations: p, penis; t, tegmen.

FIGURE 33. Scirtes sulcigerus Champion, syntype, male. A) sternites I–VII, B) tegmen and penis in ventral aspect. Abbreviations: p, penis; t, tegmen.

FIGURE 34. Scirtes crassicornis Champion, holotype, male. A) tergite VIII, B) tergite IX, C) sternite IX, D) tegmen and penis in ventral aspect.

FIGURE 36. Scirtes malayanus Champion, holotype, male. A) tergite VIII, B) tergite IX, C) sternite IX, D) tegmen and penis in ventral aspect.

FIGURE 37. Scirtes cameroni sp. n., holotype, male. A) tergite VIII, B) tergite IX, C) sternite IX, D) tegmen and penis in ventral aspect.

FIGURE 38. Scirtes sabahensis sp. n., holotype, male. A) tergite VIII, B) tergite IX, C) sternite IX, D) tegmen and penis in ventral aspect.

FIGURE 39. Scirtes brancuccii sp. n., holotype, male. A) sternites V–VII, B) tergite VIII, C) tergite IX, D) sternite IX, E) tegmen and penis in ventral aspect.