Fannia chingaza, Grisales, Diana, Wolff, Marta & De, Claudio J. B., 2012

Grisales, Diana, Wolff, Marta & De, Claudio J. B., 2012, Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia, Zootaxa 3591, pp. 1-46 : 10-11

publication ID	https://doi.org/ 10.5281/zenodo.213946
DOI	https://doi.org/10.5281/zenodo.6175045
persistent identifier	https://treatment.plazi.org/id/03BE87F8-5238-D423-C094-2419FB77FA16
treatment provided by	Plazi
scientific name	Fannia chingaza
status	sp. nov.

Treatment

Fannia chingaza View in CoL sp. nov.

( Figs. 22 View FIGURES 20 – 37 , 40 View FIGURES 38 – 55 , 58 View FIGURES 56 – 73 , 76 View FIGURES 74 – 82 , 94 View FIGURES 92 – 100 , 112 View FIGURES 110 – 118 , 128 View FIGURES 128 – 131 )

Diagnosis. these characters apply only to the male sex. Calypters whitish; fore tarsomeres 3–5 flattened and dark; hind femur weakly curved, posteroventral and posterior surfaces densely setulose, with weak, long setae with hooked apices forming a tuft up to the pre-apical protuberance, ventral surface with pre-apical protuberance extended to anterior and posterior surfaces; terminalia with spiral bacilliform process.

Description. Holotype male: body length 5.6 mm; wing length 5.5 mm.

Head: eye densely setulose. Frontal vitta velvety black, brownish pollinose. Interocular space 0.16 mm. 13-14 fr. Ocellar triangle black; oc proclinate and developed, 4 less developed setulae on ocellar area. Poc divergent. Fronto-orbital plate black, greyish pollinose. Parafacial bare, narrowing ventrally, greyish pollinose on dorsal half and yellowish on ventral half. Fronto-genal suture with brownish hairs. Gena black, yellowish pollinose. Scape black, with 3 short setae reaching pedicel. Pedicel black pollinose, dorsal setae developed. Postpedicel black, brownish pilose, 2 times the length of pedicel. Arista black, pubescent. Palpus black and filiform.

Thorax: black, weakly brownish pollinose; scutum without vittae. Acr 2:3. Dc 2:3, long. Pprn 4, 2 developed and 2 shorter. Pra 2, seta closer to the suture longer. Prepm setulose. Pre-basal scutellar area with ground setulae. Subapical scutellar setae 1 pair. Discal scutellar setae 1 pair. Lateral scutellar setae present.

Wing: brownish, with upper third between vein C and half of vein R 2+3, and cells br, bm and cup darker. Calypters whitish. Haltere brown with stem and base lighter in colour.

Legs: black with tarsi black and pulvilli brown. Fore femur with 1 row of long d; a and av inconspicuous; ventral surface bare; 1 row of long pv with straight apices; posterior surface setulose, with long and weak setae with curved apices. Fore tibia with 1 strong pre-apical d; ad on edge of apical third and pre-apical d area; 1 short apical v; 1 pre-apical pv; 1 p on edge of apical third. Fore tarsus with 1 strong and thin v on first tarsomere; tarsomeres 3–5 slightly flattened ( Fig. 128 View FIGURES 128 – 131 ). Mid femur constricted on pre-apical ventral surface; 1 row of short ad ending in 3 long pre-apicals; 1 row of strong av that decrease in length and become sparse towards apex; 1 row of v with hooked apices on basal third; 1 row of long pv with hooked apices, setae decrease in length towards apex, 4 strong setae with straight apices on basal third; 1 row of long, weak p with curved apices, 6 longer p with straight apices on apical third. Mid tibia on ventral surface with a pronounced constriction on basal and medial half and pre-basal area, dorsal surface densely setulose on apical half, 1 long and strong apical v; 1 developed pre-apical ad; 1 developed a on edge of apical third and 2 apical, one long, developed, and one short; 1 short apical v; 1 apical pv; 1 p on edge of apical third and 2 pre-apical. Hind coxa bare on posterior margin. Hind femur slightly curved; ventral surface with pre-apical protuberance extending over anterior and posterior surfaces ( Figs. 22 View FIGURES 20 – 37 , 40 View FIGURES 38 – 55 ); 1 developed d on apical third; 1 long ad on apical third; 1 row of a, starting on apical third, 3 setae running towards anterodorsal surface ( Fig. 22 View FIGURES 20 – 37 ); anteroventral surface setulose, with setae that increase in length up to halfway into pre-apical protuberance and end up in a tuft of long setae with hooked apices ( Fig. 22 View FIGURES 20 – 37 ); posteroventral and posterior surfaces densely setulose, with weak, long setae with hooked apices, forming a tuft that goes halfway into the preapical protuberance ( Fig. 40 View FIGURES 38 – 55 ). Hind tibia with 1 developed median d and 1 developed pre-apical; 1 long and developed median ad at the same level of d; 1 short apical a; 2 median av, apical most seta longer, 1 apical ( Fig. 22 View FIGURES 20 – 37 ).

Abdomen: elongate, black, weakly greyish pollinose. Syntergite 1+2 with a set of long lateral setae. Tergites 3–5 setulose laterally, with long and developed setae. Sternite 1 densely setulose. Sternite 5 as in Fig. 58 View FIGURES 56 – 73 . Terminalia ( Figs. 76 View FIGURES 74 – 82 , 94 View FIGURES 92 – 100 ): epandrium wider than long, with apical median expansions and a few weak setae on basal half; cercal plate elongate apically where it is more heavily sclerotized, with short weak setae along the plate; surstylus strongly articulated with epandrium, curved apically, shaped as a comma in lateral view, basal half with short lateral expansion, with short and weak setae on the inner side, apices of surstyli crossed; bacilliform process spiralled and simple; hypandrium and associated structures as in Fig. 112 View FIGURES 110 – 118 .

Female: unknown.

Biology. according to collecting data, Fannia chingaza sp. nov. is associated with the cold climates of the Páramo ecosystem (above 3000 m) and occurs at the Parque Nacional Natural Chingaza . The Páramo ecosystem is characterized by an abundance of water bodies, “frailejón ( Asteraceae , Espeletia ), bog mosses, lichens and species richness of the endemic fauna and flora (Parques Nacionales Naturales de Colombia 2002).

Comments. these characters apply only to the male sex. Fannia chingaza sp. nov. is morphologically similar to Fannia iguaque sp. nov. They differ on the hind femur (with a tuft of more developed setae on pre-apical protuberance in F. c h i n g a z a), and fore tarsomere (less flat in F. c h i n g a z a). The fore tarsus with modified tarsomeres, basal tarsomere on ventral surface with basal differentiated seta and spiralled bacilliform process (Pont & de Carvalho 1994) point to the placement of this species in the anthracina group. However, F. chingaza lacks the white colour on the fore tarsus.

Etymology. name in apposition. The species epithet refers to the Parque Nacional Natural Chingaza , typelocality of the species. The pre-Colombian Muisca, to whom Chim-gua-za meant “Night Mountain God, held the Páramo de Chingaza as a sacred site.

Type material. Holotype male (IAvH).CO [ Colombia]. Cundinamarca. P.N.N [Parque Nacional Natural] Chingaza \ Alto de La Bandera\ 4˚31’N 73˚45W\ 3660 m. Malaise. 3–15. III. 2001 \ L. Cifuentes y E. Raigoso\ Leg. M. 1489. IAvH. The holotype is in excellent condition and has all structures.

Distribution. Colombia: department of Cundinamarca (Andean region, Eastern Cordillera).