Pseudonortonia camarinensis, Selis & Nguyen, 2024

Selis, Marco & Nguyen, Lien Thi Phuong, 2024, The Oriental species of the genus Pseudonortonia Giordani Soika (Hymenoptera Vespidae: Eumeninae), Zootaxa 5477 (2), pp. 171-194 : 176-179

Pseudonortonia camarinensis sp. nov.

( Fig. 4 )

Diagnosis. The female of this species can be distinguished from the other Oriental species of Pseudonortonia by the following combination of characters: clypeus sparsely and finely punctate; occipital carina shallow and straight on gena; pronotal carina almost entirely absent dorsally; propodeal shelf longer than metanotum and shallowly furrowed, about at same level as scutellum; T1 1.15× as long as apically wide and densely punctate, interspaces forming reticulation; T2 with punctures becoming abruptly finer and sparser on disc, long smooth apical part not preceded by elongate pits and about 4× as long as apical lamella; S2 evenly convex.

Material examined. HOLOTYPE: ♀ labeled “P.I., CAMARINES / SUR, Mt. Iriga / 500m, 31.III.1962 // H.M. Torrevillas / Collector / BISHOP // Pseudonortonia / aff. bhamens / Giordani Soika det. 1981 // Pseudonortonia camarinensis / HOLOTYPUS ♀ / Det. Marco Selis [red label]” ( MSNVE).

Description. Female holotype. Body length 8.6 mm; fore wing length 7.0 mm.

Head in frontal view about 1.2× as wide as high. Inner eye margins converging below, interocular distance about 1.5× wider at vertex than at clypeus. Clypeus in frontal view about 1.1× as wide as high; basal margin very weakly convex, narrowly separated from antennal toruli; apical margin shallowly emarginate, 0.3× as wide as clypeus, lateral teeth apically rounded and slightly pointing out; clypeus in lateral view weakly convex, shallowly depressed near apex. Gena 0.65× as wide as eye at level of bottom of ocular sinus; distance from posterior ocellus to occipital carina about 1.65× as distance from posterior ocellus to inner eye margin; cephalic foveae very small and close to each other, placed in a barely developed smooth convex area which continues posteriorly in an irregular longitudinal carina; occipital carina complete, higher and shallowly reflexed on gena and middle of vertex, much weaker on sides of vertex, weakly and evenly curved on gena. Mandible with five teeth. Scape 3.2× as long as apically wide; F1 1.35× as long as wide, F2 subquadrate, F3–9 wider than long. Mesosoma 1.55× as long as wide. Pronotal carina strong and sharp on lateral faces, strongly produced and angled on humeri, where it forms a lobe that is triangular in frontal view and spiniform in dorsal view; pronotal carina absent on the dorsal face, except for a very weak indication in the middle. Mesoscutum 1.1× as long as wide; notauli visible in the posterior third; mesoscutum in lateral view weakly convex, somewhat flattened posteriorly. Scutellum smoothly passing into mesoscutum, entirely flattened; anterior margin crenate and forming five pits, the median one larger; median longitudinal furrow present but very shallow. Metanotum almost horizontal, very weakly sloping posteriorly and shallowly furrowed medially. Tegula broadly rounded, with anterior third of outer margin nearly straight, posterior lobe more or less right-angled and reaching apex of parategula; parategula digitiform and curved. Epicnemial carina complete, well-developed but blunt, almost reaching pronotal margin. Posterior face of propodeum with sharp median carina running from base to apex; dorsal faces of propodeum meeting along mid-line and forming a propodeal shelf, which is longer than the metanotum and shallowly furrowed; dorsal carinae present but irregular due to coarse sculpture, ventral carinae present and sharp, lateral carinae absent; submarginal carina produced into a subtriangular but apically rounded lobe above valvula; propodeal valvula with deeply emarginate posterior margin, not completely fused to submarginal carina. T1 subpetiolate and narrower than T2, in dorsal view about 1.15× as long as apically wide, sides evenly diverging from carina to apex; T 1 in lateral view with a barely developed basal petiole, then abruptly swollen with evenly convex dorsal margin, barely depressed apically; transverse carina strong on the dorsal face, distinctly angled on the sides and then shortly continued on lateral face, where it is blunt. S1 subtriangular with concave apical margin, completely separated from T1 and bearing a pair of strong but blunt carinae running from basal petiole to posterior corners. T 2 in dorsal view with weakly and evenly convex sides, barely constricted apically; in lateral view more or less evenly convex, with shallowly depressed apical margin; apical margin with a series of coarse punctures basally delimiting a long smooth area, which is 4× as long as apical hyaline lamella. S 2 in lateral view barely longer than T2 and evenly convex; apical margin similar to T2, but hyaline lamella longer. Prestigma subquadrate, about 0.20× as long as pterostigma; second recurrent vein nearly interstitial.

Clypeus with very sparse and fine punctures, interspaces equal to several puncture diameters and strongly shiny, with very sparse micropunctures. Frons with dense deep punctures, interspaces narrower than puncture diameter and forming a shiny and shallowly convex reticulation. Vertex and gena similar to frons but with sparser punctures, some interspaces reaching one puncture diameter in length; lower narrow part of gena with irregular and sparse punctures; occipital carina crenate on middle of vertex and less evidently on gena, with a short longitudinal carina in the middle. Scape with sparse micropunctures and few scattered fine macropunctures. Pronotum similar to frons but punctures larger; anterior face sculpted like dorsal faces but with larger smooth interspaces; lateral faces with traces of longitudinal striation. Mesoscutum with larger and coarser punctures, interspaces forming irregular longitudinal ridges in the area between the notauli. Scutellum similar to mesoscutum, longitudinal ridges more irregular. Metanotum with few large and coarse punctures, sides mostly smooth and shiny. Tegula strongly shiny, completely smooth on disc, with dense micropunctures on margins. Mesepisternum with punctures similar to mesoscutum, partly arranged in oblique series; epicnemium crenate along border of foramen, with deep punctures on mesosternum. Dorsal plate of metaepisternum with strong longitudinal ridges, median ridge particularly developed; ventral plate largely shiny and micropunctate, with crenate anterior margin and small deep punctures on anteroventral corner. Dorsal faces of propodeum coarsely punctate, punctures large and deep, interspaces strongly raised and forming reticulation; lateral faces with shallower and smaller flat-bottomed punctures, mostly arranged in longitudinal series; posterior face shagreened and silky-shiny, with large shallow punctures along dorsolateral margins and oblique striae covering most of the remaining surface. Post-carinal area of T1 with large and coarse punctures, slightly sparser basally, interspaces very narrow and forming a strong reticulation; pre-carinal area mostly smooth with a curved series of deep punctures in the upper half; apical swollen margin completely smooth. T2 on basal third with deep punctures becoming denser laterally, punctures becoming much finer and sparser posteriorly on disc; T2 preapically bearing a narrow area where punctures become rapidly coarser; area between coarse punctures and apical lamella completely smooth. T3–6 shagreened and with minute punctures becoming progressively sparser and less visible. S1 with coarse and deep punctures in the area between the longitudinal carinae, transversely ridged on sides. S2 with sparse fine punctures, similar to those on disc of T2; apical smooth area preceded by an irregular series of sparse larger punctures. S3–6 similar to respective tergites. Pale golden setae on whole body, longer on frons and sides of mesosoma, shorter and very sparse on tergites; short and dense pubescence on lower part of face including clypeus, gena, sides of mesosoma and posterior face of propodeum; metasoma with dense and darker pubescence, scattered setae denser on sternites than on tergites.

Black; following parts yellow: spots on basal and apical corners of clypeus fused to form an irregularly Ushaped marking, mandible except inner margin, elongate interantennal spot, lower face of scape, small spot behind upper lobe of eye, narrowly separated triangular spots on middle of pronotum, suffused spots on tegula, parategula, narrowly separated spots on metanotum, narrow apical band on T1 shortly curved on sides, whole apical area of T2 and S2, apical third of fore femur, outer face of all tibiae (except apex of hind tibia), short line on fore basitarsus; following parts yellowish-ferruginous: lower face of flagellum, tegula, part of fore tibia, most of fore tarsi. Wings fusco-hyaline, with suffused spot at apex of marginal cell.

Male. Unknown.

Distribution. Philippines: Luzon ( Fig. 10 ).

Etymology. The species name refers to the province of Camarines Sur, where this species was collected. It is an adjective.

Figures

FIGURE 4. Pseudonortonia camarinensis sp. nov., female holotype.A, habitus, dorsal view. B, habitus, lateral view. C, labels. D, head, frontal view. E, mesosoma, dorsal view. F, T1–2, dorsal view.

FIGURE 10. Distribution map of the Oriental species of Pseudonortonia.