Parachorius semsanganus Tarasov & Keith, 2011

Tarasov, Sergei, 2017, A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data, Zootaxa 4329 (2), pp. 101-149 : 139

Parachorius semsanganus Tarasov & Keith, 2011

( Figs. 22 , 27D )

Parachorius semsanganus Tarasov & Keith, 2011: 51

Type locality. Phou Sane Mountain, northern Laos.

Distribution. Northern Laos, Phou Sane Mountain.

Diagnosis. This species stands out in being the largest representative of Parachorius . It can be separated from all other species of Parachorius by metatibia with large teeth in males and inner margin denticulate and parameres with apex bent upward and membranous lobes.

Description. Body. Body black, length 8.2–10.6 mm. Elytra uniformly colored.

Head. Clypeus with two teeth, outer margin of each tooth very slightly notched basally.

Pronotum. Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1–2 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.

Elytra. Elytra covered with sparse punctures.

Wings. Wings normally developed.

Legs. Profemur with almost straight or sinuate anterior margin. Protibia with apical spur reaching protarsomere 2. First tooth notably modified in males, approximately twice as wide as second tooth; first and second teeth distinctly separated from each other; first tooth distinctly separated from denticles located between first and second teeth. Metafemoral posterior margin serrate (sometimes slightly), simple, not modified. Metatibia slightly sinuate, inner margin denticulate with large teeth on inner margin, produced with a small angle apically.

Metasternum. Transverse basal ridge of metasternum reduced and extremely short.

Aedeagus. Parameres symmetrical, bearing membranous lobes and bending upward apically.

Material examined. Holotype: Laos: 1 male, " LAOS-NE, Xieng Khouang prov., 19°38.20'N 103°20.20'E, Phonsavan (30 km NE) : PHOU SANE Mt. , 1420 m, 10.-30.v.2009, D. Hauck leg. / HMB Basel , NMPC Prague Laos 2009 Expedition : M. Brancucci, M. Geiser, Z. Kraus , D. Hauck, V. Kubán / HOLOTYPE Parachorius semsanganus D. Keith & S. Tarasov det. 2011", 19°38.2'N, 103°20.2'E ( NHMB) . Paratypes: 4 males, 3 females, Xieng Khouang Province, 30 km NE Phonsavan : Ban Na, Lam Phou Sane Mountain, 19°38'N, 103°20'E, 1300– 1500 m, on 10–30/v/2009 (M. Brancucci) ( NHMB) . Paratypes: 2 males, Xieng Khouang Province, Phonsavan (30 km NE): Phou Sane Mountain , Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán, 19°38.2'N, 103°20.2'E, 1420 m, on 10–30/v/2009 (Z. Kraus) ( NHMB) . Paratypes: 9 females, Xieng Khouang Province, Phonsavan (30 km NE): Phou Sane Mountain , Laos 2009 Expedition: M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubán, 19°38.2'N, 103°20.2'E, 1420 m, on 10–30/v/2009 (D. Hauck) ( NHMB) .

Figures

FIGURE 22. Morphology of Parachorius semsanganus. (A) aedeagus, lateral view (paratype). (B) aedeagus, dorsal view (paratype). (C) aedeagus, apical view (paratype). (D) aedeagus lateral view. (E) right metafemur (male, paratype). (F) habitus (male, holotype).

FIGURE 27. Distribution maps of Parachorius species. (A) Parachorius peninsularis. (B) Parachorius pseudojavanus. (C) Parachorius schuelkei. (D) Parachorius semsanganus. (E) Parachorius solodovnikovi. (F) Parachorius thomsoni. Type locality — type locality of species (holotype or lectotype); Type. Collection data — localities of type specimens associated with species (paratypes or paralectotypes); Collection data — localities of other specimens examined in this study. Literature data — localities retrieved from literature.