Parachorius nudus ( Sharp, 1875 ) S. Tarasov, 2017

Tarasov, Sergei, 2017, A cybertaxonomic revision of the new dung beetle tribe Parachoriini (Coleoptera: Scarabaeidae: Scarabaeinae) and its phylogenetic assessment using molecular and morphological data, Zootaxa 4329 (2), pp. 101-149 : 133-134

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Parachorius nudus ( Sharp, 1875 )

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Parachorius nudus ( Sharp, 1875)   new combination

( Figs. 18 View FIGURE 18 )

Cassolus nudus Sharp, 1875:40  

Cassolus nudus   ; Gillet 1911: 39

Cassolus nudus   ; Boucomont & Gillet 1921: 4 Cassolus nudus   ; Paulian 1945: 58

Panelus quadridentatus Balthasar, 1952: 223   (species inquirenda) Cassolus quadridentatus   ; Balthasar 1963: 260 Cassolus nudus   ; Balthasar 1963: 260

Cassolus nudus   ; Masumoto, 1987: 127

Cassolus nudus   ; Hanboonsong & Masumoto 2001: 135 Cassolus nudus   ; Masumoto et al. 2012: 105

Type locality. Cambodia ( Cassolus nudus   ); Hoa-Binh   , Vietnam ( Panelus quadridentatus   )

Distribution. The species is known only from the holotype specimen collected in Cambodia, which lacks any detailed information on the type locality.

Diagnosis and taxonomic notes. In external morphology this species is most similar to P. solodovnikovi   , P. longipenis   , and P. peninsularis   . Reliable species identification in this group requires examination of the parameres. The squeezed aedeagus of the holotype, the only available specimen of this species, does not allow reconstructing the real shape of the aedeagus. This imposes problems on the identity of this species and its separation from the similar species. Although the aedeagus is squeezed, the examination suggests that its parameres seem to lack hornlike dorsal lobes that are present in P. solodovnikovi   and P. longipenis   . Given current state of knowledge, this indicates that P. nudus   is a separate species. At the same time, due to squeezing the separation of the paramere shape between P. nudus   and P. peninsularis   does not seem possible. However, from the latter, P. nudus   can be reliably distinguished by the modified first tooth of the protibia, where P. nudus   is bifurcate, spatulate, bent downward, and notably larger than the second tooth with which the first tooth is almost merged. These facts support the species identity of P. nudus   .

Panelus quadridentatus   , the synonym of P. nudus   , is described from Vietnam (“ Tonkin: Hoa-Binh, A. de Cooman leg.”). All studied material from Vietnam suggests that P. nudus   does not likely occur there, which points out that P. quadridentatus   is probably conspecific with either P. solodovnikovi   or P. longipenis   , which are known from the same collecting events. Unfortunately, the type of Panelus quadridentatus   deposited in NMPC is probably lost ( Bezdek & Hajek 2011). Moreover, the original description of Panelus quadridentatus   does not list any characters enabling reliable separation of this species from other Parachorius   . Hence, being unable to examine the type and separate P. quadridentatus   from other Parachorius   , I follow Balthasar (1963) and tentatively treat P. quadridentatus   as a synonym of P. nudus   . The elucidation of P. quadridentatus   identity needs further investigation and this species has to be formally regarded as species inquirenda.

The type of P. nudus   is the only Parachorius   known from Cambodia. Additional sample of Parachorius   from Cambodia is needed to assess diagnosis and identity of P. nudus   .

Description. Body. Body dark, length 4.8 mm. Elytra uniformly colored.

Head. Clypeus with two teeth, outer margin of each tooth distinctly notched basally.

Pronotum. Pronotum rounded laterally. Frontal angles rounded. Disc covered with punctures separated by 1.0– 1.5 puncture diameter(s). Pronotum in anterior-posterior direction notably shorter than elytra.

Elytra. Elytra covered with sparse punctures.

Wings. Wings normally developed.

Legs. Profemur with almost straight or sinuate anterior margin. Protibia with apical spur reaching protarsomere 2. First tooth notably modified in males, spatulate, bent downward and notably larger than second tooth, with which it almost merged; first tooth may look bifurcated as it is merged with denticles located between first and second teeth. Metafemoral posterior margin not serrate, simple, not modified. Metatibia slightly sinuate, inner margin denticulate with relatively small teeth, produced in small angle apically.

Metasternum. Transverse basal ridge of metasternum reduced and extremely short.

Aedeagus. Parameres symmetrical, simple, and narrowed toward apex (lateral view).

Material examined. Holotype of Cassolus nudus   : Cambodia: 1 male, " Cassolus nudus   type DS / Camb Mouhot [ Cambodia] / Ex Musaeo D.Sharp 1890 / G..J. Arrow vidit 1901 / Museum Paris ex. coll. R. Oberthur / Type / Holotypus Cassolus nudus S. Tarasov   det. 2010 / Parachorius nudus S. Tarasov   det. 2011 " ( MNHN).


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Parachorius nudus ( Sharp, 1875 )

Tarasov, Sergei 2017

Cassolus nudus

Masumoto 2012: 105
Hanboonsong 2001: 135

Cassolus nudus

Masumoto 1987: 127

Panelus quadridentatus

Balthasar 1963: 260
Balthasar 1963: 260
Balthasar 1952: 223

Cassolus nudus

Paulian 1945: 58
Boucomont 1921: 4

Cassolus nudus

Gillet 1911: 39

Cassolus nudus

Sharp 1875: 40