Philonthus pulcher, Wendeler, 1924
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publication ID |
https://doi.org/ 10.1206/0003-0090.265.1.5 |
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persistent identifier |
https://treatment.plazi.org/id/03C1F679-FEC6-FECF-FF2F-FF7B8392F920 |
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treatment provided by |
Felipe |
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scientific name |
Philonthus pulcher |
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pulcher Wendeler, 1924 View in CoL , see: quadripunctatus Bernhauer, 1915 .
quadripunctatus Bernhauer, 1915 g: 193 ( Leucitus ; Type locality: Neu-Guinea: Sattelberg). — Scheerpeltz, 1933: 1380 ( Leucitus ; catalog).
— Cameron, 1937a: 104 ( Leucitus ; New Guinea).
— Last, 1975a: 1 ( Leucitus ).
— Last, 1975a: 3 ( Leucitus ; characters).
— Last, 1987: 55 ( Leucitus ; Papua New Guinea; Irian).
- DISTRIBUTION: New Guinea.
pulcher Wendeler, 1924: 340 ( Leucitus ; Type locality: Neu-Pommern, Süd-See).
— Scheerpeltz, 1933: 1380 ( Leucitus ; catalog).
— Last, 1975a: 1 ( Leucitus ; synonym of quadripunctatus ).
rufipennis Cameron, 1918 , see: Hesperus .
semichalceus Heller, 1910: 9 ( Leucitus ; Type locality: Papua New Guinea, 780–900 m).
— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).
— Last, 1975a: 5 ( Leucitus ; characters).
- DISTRIBUTION: New Guinea.
somoruensis Last, 1968: 360 ( Philonthus ; Type locality: New Guinea: Mobutie Somoru).
— Last, 1975a: 1, 4 ( Leucitus ; characters).
- DISTRIBUTION: New Guinea.
stenoides Gravenhorst, 1806: 90 ( Staphylinus ; Type locality: Iauae).
— Erichson, 1840: 493, 930 ( Philonthus ; Java).
— Fauvel, 1878f: 253 ( Leucitus ; characters; Java).
— Bernhauer and Schubert, 1914: 373 ( Leucitus ; catalog).
— Last, 1975a: 1 ( Leucitus ).
— Schillhammer, 1999b: 95 ( Leucitus ; based on original description, species definitely belongs in Leucitus ; type locality in Java probably erroneous; species probably from New Guinea; type specimen probably lost).
- DISTRIBUTION: Indonesia.
superbus Arrow, 1915: 499 ( Philonthus ; Type locality: Utakwa, River, Dutch New Guinea). — Scheerpeltz, 1933: 1363 ( Philonthus ; catalog).
— Last, 1975a: 1, 6 ( Leucitus ; characters).
- DISTRIBUTION: New Guinea.
[1 species; Neotropical Region]
Linoderus Sharp, 1885: 452 (species included: gracilipes ). TYPE SPECIES: Linoderus gracilipes Sharp , fixed by monotypy.
— Bernhauer and Schubert, 1914: 367 (world catalog; 1 species).
— Blackwelder, 1944: 136 (checklist of species from Mexico, Central and South America, and the West Indies).
— Blackwelder, 1952: 221 (type species: gracilipes ).
gracilipes Sharp, 1885: 452 ( Linoderus ; Type locality: Panama, Volcan de Chiriqui, Bugaba). — Bernhauer and Schubert, 1914: 367 ( Linoderus ; catalog).
- DISTRIBUTION: Panama.
[8 species; Ethiopian Region]
Mentophilonthus Levasseur, 1966: 207 (species included: odzalaensis , triseriatus , descarpentriesi , mongendensis , dilutior , lampropterus ). TYPE SPECIES: Philonthus triseriatus Bernhauer , fixed by original designation.
bangoranensis Levasseur, 1980: 354 ( Mentophilonthus ; Type locality: République
Centrafricaine: Bangoran, SW de Ndélé).
centrafricanus Levasseur, 1980: 356 ( Mentophilonthus ; Type locality: République Centrafricaine: Bangoran , SW de Ndélé).
- DISTRIBUTION: Central African Republic.
descarpentriesi Levasseur, 1966: 208 , 209 ( Mentophilonthus ; Type locality: Congo ex français, Odzala. Also cited from Belgian Congo).
— Cameron, 1929: 61 ( Philonthus ; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).
— Tottenham, 1949b: 300 ( Philonthus ; [Note: Misidentified as triseriatus according to Levasseur, 1966: 209]; characters; Belgian Congo).
— Levasseur, 1966a: 1211 ( Mentophilonthus ; Belgian Congo; Congo).
- DISTRIBUTION: Congo Republic, Congo.
dilutior Bernhauer, 1928 g: 113 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1340 ( Philonthus ; catalog).
— Levasseur, 1966: 208, 211 ( Mentophilonthus ; characters; Belgian Congo).
- DISTRIBUTION: Congo.
seriatipennis Bernhauer, 1928 g: 114 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).
— Cameron, 1929: 61 ( Philonthus ; characters; Belgian Congo).
— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).
— Tottenham, 1949b: 300 ( Philonthus ; characters).
— Levasseur, 1966: 211 ( Mentophilonthus ; synonym of dilutior ).
lampropterus Bernhauer, 1934 g: 239 ( Philonthus ; Type locality: Kivu: Kibumba ).
— Cameron, 1950g: 35 ( Philonthus ; Belgian Congo).
— Levasseur, 1966: 213 ( Mentophilonthus ; characters; Belgian Congo).
- DISTRIBUTION: Congo.
minor Bernhauer, 1928 , see: triseriatus Bernhauer, 1928 .
mongendensis Bernhauer, 1928 g: 114 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende). — Scheerpeltz, 1933: 1351 ( Philonthus ; catalog).
— Levasseur, 1966: 208, 211 ( Mentophilonthus ; characters; Belgian Congo).
- DISTRIBUTION: Congo.
odzalaensis Levasseur, 1966: 208 , 211 ( Mentophilonthus ; Type locality: Odzala , Congo ex française. Also cited from Gabon).
— Levasseur, 1966a: 1211 ( Mentophilonthus ; Congo; Zaire; Gabon).
— Levasseur, 1968a: 53 ( Mentophilonthus ; Central African Republic).
- DISTRIBUTION: Gabon, Congo Republic, Congo, Central African Republic.
parcior Bernhauer, 1928 , see: triseriatus Bernhauer, 1928 .
seriatipennis Bernhauer, 1928 , see: dilutior Bernhauer, 1928 .
triseriatus Bernhauer, 1928 g: 112 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1365 ( Philonthus ; catalog).
— Levasseur, 1966: 208 ( Mentophilonthus ; characters; Congo; Ruanda).
— Levasseur, 1966a: 1210 ( Mentophilonthus ; Congo; Zaire).
- DISTRIBUTION: Congo Republic, Congo, Rwanda.
minor Bernhauer, 1928 g: 113 ( Philonthus ; [preoccupied]; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1365 ( Philonthus ; variety of triseriatus ).
parcior Bernhauer, 1928 g: 113 ( Philonthus ; Type locality: Belgisch-Kongo: Mongende).
— Scheerpeltz, 1933: 1365 ( Philonthus ; variety of triseriatus ).
[13 species; Ethiopian Region]
Moeocerus Fauvel, 1899: 100 (replacement name for Homoeocerus Fauvel ). TYPE SPECIES:
— Bernhauer and Schubert, 1916: 410 (world catalog; 7 species).
— Scheerpeltz, 1933: 1420 (world catalog supplement).
— Gridelli, 1939: 322 (characters; key to species).
Homoeocerus Fauvel, 1899a: 27 (species included: mimus , simplex , ciliatus , robustus , spinulosus , puncticollis ; preoccupied by Burmeister, 1835, and Kolenati, 1859). TYPE SPECIES: Homoeocerus mimus Fauvel by objective synonym with Moeocerus for which Blackwelder fixed the type species. [Note: Blackwelder stated that R. Lucas, 1920: 421, designated the type species but the designation was ambiguous].
— Fauvel, 1907: 48 (synonym of Moeocerus ).
— Bernhauer and Schubert, 1916: 410 (synonym of Moeocerus ).
— Blackwelder, 1952: 190 (type species; synonym of Moeocerus ).
Polystictomoeocerus Scheerpeltz, 1977: 145 (subgenus of Moeocerus ; species included: puncticollis , griseosericans ). TYPE SPECIES: Homoeocerus puncticollis Kraatz , fixed by original designation.
abessinus Bernhauer, 1917 g: 46 ( Moeocerus ; Type locality: Abessinien: Harrar ).
— Scheerpeltz, 1933: 1420 ( Moeocerus ; catalog).
— Gridelli, 1939: 326, 329 ( Moeocerus ; characters; Abyssinia).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Ethiopia.
ciliatus Roth, 1851: 118 ( Acylophorus ; Type locality: Abyssinien).
— Fauvel, 1899a: 30 ( Homoeocerus ; characters; Tigre; Abyssinie).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Gridelli, 1939: 326 ( Moeocerus ; characters; Abyssinia).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Ethiopia.
doriae Gridelli, 1939: 335 ( Moeocerus ; Type locality: Cheren , Eritrea; regione dell’Eritrea, Paese dei Bogos).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Eritrea.
fauveli Gridelli, 1939: 337 ( Moeocerus ; Type locality: Maritzburg, Natal).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: South Africa.
griseosericans Scheerpeltz, 1977: 143 ( Moeocerus ; subgenus Polystictomoeocerus ; Type locality: Ed. Luja in der Umgebung von Kondué im Kongogebiet).
- DISTRIBUTION: Congo.
lineiventris Fauvel, 1907: 48 ( Moeocerus ; Type locality: Kilimandjaro: Kibosho , 1700 m).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Tanzania.
mimus Fauvel, 1899a: 29 ( Homoeocerus ; Type locality: Natal).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Gridelli, 1939: 326, 334 ( Moeocerus ; characters).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: South Africa.
overlaeti Bernhauer, 1935d: 104 ( Moeocerus ; Type locality: Lulua: Kapanga).
— Gridelli, 1939: 326 ( Moeocerus ; Abyssinia).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Congo, Ethiopia.
puncticollis Kraatz, 1858: 363 ( Homorocerus ; Type locality: Senegambia).
— Fauvel, 1899a: 32 ( Homoeocerus ; Senegambie portugaise).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Tottenham, 1961: 205 ( Moeocerus ; Senegal).
— Scheerpeltz, 1977: 145 ( Moeocerus ; subgenus Polystictomoeocerus ).
- DISTRIBUTION: “Senegambia”, Senegal, Ethiopia.
robustus Gestro, 1881: 658 ( Homorocerus ; Type locality: Let Marefia).
— Fauvel, 1899a: 30 ( Homoeocerus ; characters; Abyssinie; Choa: Let Marefia; Giuba).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Gridelli, 1939: 330 ( Moeocerus ; characters; Abyssinia).
— Scheerpeltz, 1977: 146 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Ethiopia.
simplex Fauvel, 1899a: 30 ( Homoeocerus ; Type locality: Angola).
— Bernhauer, 1915o: 363 ( Moeocerus ; Belgian Congo).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Angola, Congo.
spinulosus Solsky, 1864a: 375 ( Homorocerus ; Type locality: Insel Gorea an der Westküste von Afrika).
— Fauvel, 1899a: 31 ( Homoeocerus ; characters; Natal; Abyssinia).
— Bernhauer and Schubert, 1916: 411 ( Moeocerus ; catalog).
— Bernhauer and Chapman, 1939: 73 ( Moeocerus ; Kenya).
— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Senegal, Ethiopia, Kenya, South Africa.
zavattarii Gridelli, 1939: 332 ( Moeocerus ; Type locality: Moyale; Arero).
— Scheerpeltz, 1977: 147 ( Moeocerus ; subgenus Moeocerus ).
- DISTRIBUTION: Ethiopia.
[3 species; Oriental and Australian Regions]
Mysolius Fauvel, 1878f: 255 (species included: aurichalceus ). TYPE SPECIES: Mysolius aurichlaceus Fauvel , fixed by monotypy.
— Duvivier, 1883: 152 (catalog).
— Olliff, 1887: 497 (characters; Australian species).
— Bernhauer and Schubert, 1914: 373 (world catalog; 2 species).
— Blackwelder, 1952: 256 (type species: aurichalceus ).
aurichalceus Fauvel, 1878f: 256 ( Mysolius ; Type locality: Nouvelle-Guinée, Romoi;
Iles Arou, Wokan; Mysol; Salvatti).
— Fauvel, 1879: 97 ( Mysolius ; characters; Fly River).
— Bernhauer and Schubert, 1914: 373 ( Mysolius ; catalog).
— Cameron, 1924: 87 ( Philonthus ; New Guinea).
— Cameron, 1931: 363 ( Mysolius ; New Guinea).
— Last, 1987: 55 ( Mysolius ; Irian).
- DISTRIBUTION: New Guinea, Indonesia.
diversipes Fauvel, 1879: 98 ( Mysolius ; race of aurichalceus ; Type locality: Iles Arou; Nouvelle Guinée; Fly River).
— Bernhauer and Schubert, 1914: 373 ( Mysolius ; variety of aurichalceus ).
chalcopterus Olliff, 1887: 497 ( Mysolius ; Type locality: Mulgrave River, North Queensland). — Bernhauer and Schubert, 1914: 373 ( Mysolius ; catalog).
— Bernhauer, 1920: 16 ( Mysolius ; Australia).
— Last, 1987: 55 ( Mysolius ; Papua New Guinea).
- DISTRIBUTION: Australia, New Guinea.
cupreosplendens Cameron, 1943a: 344 ( Actinus ; Type locality: New Guinea: Milne Bay).
— Last, 1987: 55 ( Mysolius ; Papua New Guinea).
- DISTRIBUTION: New Guinea.
[72 species; Neotropical, Nearctic, Palaearctic, Ethiopian,
Madagascan, Oriental, and Australian Regions]
Neobisnius Ganglbauer, 1895: 464 (species included: villosulus ; procerulus ; prolixus ). TYPE SPECIES: Gabrius villosulus Stephens , fixed by subsequent designation by R. Lucas, 1920: 434.
— Thomson, 1859: 25 (cited as Bisnius ; [Note: Thomson’s use of Bisnius was a misidentification of Neobisnius ; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters; type species: cephalotes ).
— Thomson, 1860: 168 (cited as Bisnius ; [Note: Thomson’s use of Bisnius was a misidentification of Neobisnius ; Tottenham, 1949 cited it with a type species and as a synonym of Neobisnius as though it were an available name]; characters).
— Heyden, 1880: 73 (cited as Actobius ; list of species of Siberian region).
— Heyden, Reitter, and Weise, 1906: 163 (list of species of Europe and Caucasus).
— Reitter, 1909: 116, 123 (characters; key to species).
— Petri, 1912: 71 (list of species of Siebenbürgen).
— Bernhauer and Schubert, 1914: 322 (world catalog; 39 species).
— Leng, 1920: 106 (catalog of North American species).
— Winkler, 1925: 374 (catalog for Palaearctic region).
— Cameron, 1925: 54 (catalog of Indian species).
— Porta, 1926: 99 (characters; key to species of Italy).
— Bruch, 1928a: 427 (catalog; Argentina).
— Portevin, 1929: 356 (key to French species).
— Roubal, 1930: 366 (catalog; Slovakia).
— Cameron, 1932: 56 (characters; key to species of British India).
— Joy, 1932: 126 (key to British species).
— Scheerpeltz, 1933: 1326 (world catalog supplement).
— Normand, 1935: 370 (list of species of Tunisia).
— Blackwelder, 1943: 440 (synonym of Erichsonius ; type species: cinerascens ).
— Gridelli, 1943: 111 (characters; notes; key to a few species).
— Kloet and Hincks, 1945: 165 (list of British species).
— Last, 1948: 148 (key to European species).
— Tottenham, 1949: 371, 417 (type species: villosulus Stephens = elongatulus Erichson ; checklist of British species).
— V. Hansen, 1952: 29 (characters; key to Danish species).
— Blackwelder, 1952: 258 (type species).
— Hatch, 1957: 173, 174 (characters; key to species of Pacific Northwest).
— Kocher, 1958: 142 (checklist of species; Morocco).
— Smetana, 1958: 134 (characters; habitus of one species; type species: villosulus ; key to Czechoslovakian species).
— Smetana, 1959a: 209 (list of species collected in Albania).
— Ferreira, 1962a: 17 (list of Portuguese species).
— Coiffait, 1963: 8, 18 (characters; key to French species groups and included French species).
— Székessy, 1963: 53 (characters).
— Palm, 1963b: 58 (characters; key to species of Sweden).
— Lohse, 1964: 171 (characters; key to central European species).
— Moore, 1965: 98, 100 (characters; notes).
— Coiffait and Saiz, 1968: 348, 360 (characters; notes).
— Scheerpeltz, 1968a: 64 (catalog of Austrian species).
— Shibata, 1973c: 56 (checklist of species of Taiwan).
— Tikhomirova, 1973a: 182 (checklist of species of USSR).
— Coiffait, 1974: 322 (characters; key to species of western Palaearctic region).
— Moore and Legner, 1975: 18 (catalog; North America north of Mexico).
— Pope, 1977: 29 (list of British species).
— Topp, 1978: 312 (larval characters).
— Muona, 1979: 14 (list of Scandinavian species).
— Uhlig, Vogel, and Sieber, 1980: 246 (some species collected in Germany).
— Burakowski, Mroczkowski, and Stefańska, 1980: 29 (catalog; Poland).
— Frank, 1981: 2 (revision of and key to New World species; characters; checklist of New World species; species group classification).
— Uhlig and Vogler, 1981: 110 (list of some species collected in Germany; habitat and collecting notes).
— Shibata, 1983: 88 (checklist of species of Japan).
— Hammond, 1984: 206 (checklist of species of Borneo).
— Tóth, 1984: 34 (characters; habitus; key to species of Hungary).
— Frank, 1986a: 370 (checklist of species of Florida).
— Segers, 1986: 46 (checklist of species; Belgium).
— Drugmand 1987b (comments on distribution of Belgian species).
— Campbell and Davies, 1991: 117 (checklist of species; Canada).
— Silfverberg, 1992: 17 (list of species of Fennoscandia, Denmark, and the Baltic States).
— Smetana, 1993b: 47 (list of species of Czech Republic and Slovakia).
— Hernández, Outerelo, and Gamarra, 1994: 188 (checklist of species; Canary Islands).
— Smetana, 1995e: 50 (characters).
— Ciceroni and Zanetti, 1995: 27 (list of species of Italy).
— M. Hansen, 1996: 104 (list of Danish species).
— M. Hansen, Mahler, Palm, and Pedersen, 1996: 243 (additions to list of Danish species).
— Cho, 1996a: 159 (characters; key to species of Korea).
— Downie and Arnett, 1996: 374 (characters; key to species of Northeastern North America).
— Lundgren, 1998: 51 (list of species from Florida; records for other States listed for each species).
— Bain, 1998: 39 (17th century remains; Massachusetts).
— M. Hansen, Palm, Pedersen, and Runge, 1998: 73 (list of species collected in 1997; Denmark).
— Newton, Thayer, Ashe, and Chandler, 2000: 396 (14 Nearctic species; notes; characters in key).
— Hayashi, 2000: 23 (notes on empodial setae).
adustus Sharp, 1887 , see: terminalis LeConte, 1863 .
afghanicus Coiffait, 1979d: 559 ( Neobisnius ; Type locality: entre Camdech et Kouchtous, 1400 à 2000 m).
— Tronquet, 1981: 71 ( Neobisnius ; Afghanistan).
— Coiffait, 1982b: 76 ( Neobisnius ; note).
- DISTRIBUTION: Afghanistan.
agnatus Erichson, 1840 , see: sobrinus Erichson, 1840 .
alaiensis Bernhauer, 1914b: 66 ( Neobisnius ; Type locality: Nord-Alai: Fergana; [Note: See lectotype designation by Gusarov, 1995a]).
— Scheerpeltz, 1933: 1326 ( Neobisnius ; catalog).
— Coiffait, 1974: 329 ( Neobisnius ; characters; Transcaspian region; Afghanistan).
— Gusarov, 1995a: 92 [= 1996: 55] ( Neobisnius ; lectotype designation).
- DISTRIBUTION: Uzbekistan, Afghanistan.
alternans Sharp, 1885 , see: jocosus Horn, 1884 .
angusticollis Kolenati, 1846 , see: prolixus Erichson, 1840 .
angustus Cho, 1996a: 162 ( Neobisnius ; Type locality: Korea: Chejudo , Kyoraeri).
- DISTRIBUTION: Korea.
arduus Sharp, 1885 , see: sobrinus Erichson, 1840 .
armuellensis Bierig, 1933: 56 ( Neobisnius ; Type locality: Panama: Puerti Armuelles ).
— Frank, 1981: 28 ( Neobisnius ; characters; Costa Rica; Panama; Colombia).
- DISTRIBUTION: Colombia, Panama, Costa Rica.
bakeri Bernhauer, 1916 h: 424 ( Neobisnius ; Type locality: Luzon : Mt. Makiling).
— Scheerpeltz, 1933: 1326 ( Neobisnius ; catalog).
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