Philonthus phaeopus, Stephens, 1833

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. V. Staphylinine Group (Part 2) Staphylininae: Diochini, Maorothiini, Othiini, Platyprosopini, Staphylinini (Amblyopinina, Anisolinina, Hyptiomina, Philonthina), Bulletin of the American Museum of Natural History 2001 (265), pp. 2441-3020 : 2441-3020

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.5

persistent identifier

https://treatment.plazi.org/id/03C1F679-FF19-FF1D-FF2F-FF7B8388F927

treatment provided by

Felipe

scientific name

Philonthus phaeopus
status

 

phaeopus Stephens, 1833 View in CoL , see: nigritulus Gravenhorst, 1802 .

philo Smetana, 1984: 140 ( Gabrius ; Type locality: Japan: Nikko N. P., Ryuzu).

- DISTRIBUTION: Japan.

philomimus Schillhammer, 2000 c: 125 View in CoL ( Gabrius View in CoL ; Type locality: Russia: Primorskiy Kray: Sikhote-Alin   GoogleMaps , Biol. Stat., 30 km SE Chuguyevka, 44.05N, 134.12E, 650 m).

- DISTRIBUTION: Russia.

philonthoides Schillhammer, 1992: 64 ( Gabrius ; Type locality: China: Gansu, Meijishan ).

- DISTRIBUTION: China.

picipennis Mäklin, 1852: 313 ( Philonthus ; [preoccupied]; Type locality: insula Sitkha).

— Fauvel, 1874: 237 [= 1874a: 463] ( Philonthus ; synonym of quisquiliarius ).

— Ganglbauer, 1895: 460 ( Philonthus ; synonym of nigritulus ).

— Bernhauer and Schubert, 1914: 350 ( Philonthus ; catalog).

— Hatch, 1957: 199 ( Philonthus ; subgenus Gabrius ; characters; British Columbia; Washington; Oregon).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1965: 29 ( Gabrius ; characters; notes; Labrador; St. Pierre; Newfoundland).

— Smetana, 1990: 178 ( Gabrius ; characters).

— Smetana, 1995e: 629 ( Gabrius ; characters; notes; all Provinces and Territories of Canada except Prince Edward Island; Alaska; Arizona; California; Colorado; Connecticut; Idaho; Illinois; Maine; Massachusetts; Michigan; Montana; Nebraska; Nevada; New Hampshire; New Mexico; New Mexico; New York; North Dakota; Oregon; Pennsylvania; South Dakota; Utah; Washington; Wisconsin; Wyoming).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Canada, USA.

decipiens Horn, 1884: 217 ( Philonthus ; [preoccupied]; Type locality: California).

— Bernhauer and Schubert, 1914: 334 ( Philonthus ; catalog).

— Fall, 1926: 150 ( Philonthus ; Alaska).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 629 ( Gabrius ; synonym of picipennis ).

lopezi Hatch, 1957: 199 ( Philonthus ; subgenus Gabrius ; Type locality: Washington:

Lopez Is. Also cited from British Columbia, Oregon, and Idaho).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1965: 57 ( Gabrius ; synonym of decipiens ).

— Smetana, 1995e: 629 ( Gabrius ; synonym of picipennis ).

piger Schillhammer, 1997: 32 ( Gabrius ; Type locality: NW-India [Kashmir], Cachmire, Pahalgam Kolakoi, 3400 m).

— Schillhammer, 2000 c: 128 ( Gabrius ; India).

- DISTRIBUTION: India.

piliger Mulsant and Rey, 1876: 553 [= 1877: 409] ( Gabrius ; Type locality: Corte Saint- François, en Corse).

— Bernhauer and Schubert, 1914: 347 ( Philonthus ; synonym of nigritulus ).

— Cameron, 1932: 147 ( Philonthus ; synonym of nigritulus ).

— Scheerpeltz, 1933: 1353 ( Philonthus ; synonym of nigritulus ).

— Coiffait, 1951: 110, 114 ( Gabrius ; characters in key; France).

— Smetana, 1957: 63, 68 ( Gabrius ; characters; England; Scandinavia; Italy; Corsica; Caucasus; Turkey; Germany; Czechoslovakia; Austria).

— Smetana, 1958: 247 ( Gabrius ; characters; ecology; Czechoslovakia).

— Smetana, 1960c: 311 ( Gabrius ; characters; Turkey; Sweden; Denmark; Norway; Germany; Holland; Belgium; England; France; Italy; Corsica; Austria; Czechoslovakia; Yugoslovia; Siberia ; Caucasus).

— Palm, 1963b: 106 ( Philonthus ; subgenus Gabrius ; characters; Sweden).

— Horion, 1965: 190 ( Gabrius ; Europe).

— Coiffait, 1974: 82 ( Gabrius ; characters; Europe; Asia Minor; Caucasus; Daghestan; Sweden; Britain; Italy; France; Corsica).

— Pope, 1977: 30 ( Gabrius ; Britain).

— Petrenko, 1980: 82 ( Gabrius ; Caucasus).

— Tóth, 1984: 87 ( Gabrius ; characters; Hungary).

— Lucht, 1987: 106 ( Gabrius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; Benelux; Sweden).

— Welch, 1993: 229 ( Gabrius ; ovariole number and ovary structure).

— Smetana, 1993b: 49 ( Gabrius ; Slovakia).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; Italy).

— Gerend and Braunert, 1997: 198 ( Gabrius ; notes; Luxembourg).

- DISTRIBUTION: Europe, Russia, Caucasus, Turkey.

stipes Sharp, 1910: 129 ( Gabrius ; Type locality: Anglia: Plymouth; Mickleham; Cambridge ).

— Bernhauer and Schubert, 1914: 356 ( Philonthus ; catalog).

— Gridelli, 1920: 152 ( Gabrius ; characters; England; Daghestan; Caucasus).

— Sainte-Claire Deville, 1926: 119 ( Philonthus ; subgenus Gabrius ; notes; Corsica).

— Scheerpeltz, 1933: 1362 ( Philonthus ; catalog).

— Wüsthoff, 1937: 32 ( Philonthus ; illustration of aedeagus).

— Méquignon, 1944: 20 ( Gabrius ; synonym of piliger ).

— Fagel, 1945: 40 ( Gabrius ; characters; Belgium).

— Smetana, 1952b: 119 ( Gabrius ; characters; Czechoslovakia).

— Smetana, 1953: 121 ( Gabrius ; Turkey).

— Smetana, 1953a: 178 ( Gabrius ; Savinian Alps).

— Smetana, 1957: 63 ( Gabrius ; synonym of piliger ).

— Smetana, 1958: 247 ( Gabrius ; synonym of piliger ).

— Smetana, 1960c: 311 ( Gabrius ; synonym of piliger ).

— Lohse, 1964: 189 ( Gabrius ; synonym of piliger ).

— Pope, 1977: 30 ( Gabrius ; synonym of piliger ).

pilipes Bierig, 1940 , see: chiriquiensis Herman, 2001 .

pisciformis Fauvel, 1874: 242 [= 1874a: 468] ( Philonthus ; Type locality: Var, Entraigues).

— Fauvel, 1874: 340 [= 1874b: 46] ( Philonthus ; Spain).

— Mulsant and Rey, 1876: 542 [= 1877: 398] ( Gabrius ; characters; France).

— Eppelsheim, 1887b: 432 ( Philonthus ; synonym of femoralis ).

— Ganglbauer, 1895: 459 ( Philonthus ; subgenus Gabrius ; characters; France; Spain).

— Bernhauer, 1899: 24 ( Philonthus ; notes; Siebenbürgen; Hungary; Bosnia; Austria).

— Bernhauer and Schubert, 1914: 337 ( Philonthus ; synonym of femoralis ).

— Portevin, 1929: 365 ( Philonthus ; characters; France).

— Scheerpeltz, 1933: 1342 ( Philonthus ; catalog).

— Smetana, 1957: 58 ( Gabrius ; synonym of femoralis ).

— Smetana, 1958: 236 ( Gabrius ; synonym of femoralis ).

— Smetana, 1961: 37 ( Gabrius ; synonym of femoralis ).

— Smetana, 1967b: 156 ( Gabrius ; Spain).

— Coiffait, 1974: 67 ( Gabrius ; characters; Spain; France; Pyrenees).

- DISTRIBUTION: France, Spain.

tibialis Mulsant and Rey, 1876: 540 [= 1877: 396] ( Gabrius ; Type locality: environs de Fréjus et dans le Languedoc).

— Duvivier, 1883: 148 ( Philonthus ; catalog; synonym of pisciformis ).

— Ganglbauer, 1895: 459 ( Philonthus ; synonym of pisciformis ).

— Bernhauer and Schubert, 1914: 337 ( Philonthus ; synonym of femoralis ).

— Coiffait, 1951: 105, 114 ( Gabrius ; cited as valid species; characters in key; France).

— Smetana, 1957: 59 ( Gabrius ; cited as valid species; characters; France; Spain).

— Fagel, 1959a: 96 ( Gabrius ; notes; Spain). — Smetana, 1967b: 156 ( Gabrius ; synonym of pisciformis ).

— Coiffait, 1974: 67 ( Gabrius ; synonym of pisciformis ).

gavoyi Bernhauer, 1928b: 12 ( Philonthus ; subgenus Gabrius ; Type locality: Gavoy in Südfrankreich; Granada in Spanien).

— Scheerpeltz, 1933: 1344 ( Philonthus ; catalog).

— Smetana, 1967b: 156 ( Gabrius ; synonym of pisciformis ).

— Smetana, 1973b: 126 ( Gabrius ; notes; synonym of pisciformis ).

— Coiffait, 1974: 67 ( Gabrius ; synonym of pisciformis ).

plumosus Last, 1981: 125 ( Gabrius ; Type locality: Marifuanga, Asaro-Chimbu Divide).

— Last, 1987: 55 ( Gabrius ; Papua New Guinea).

— Schillhammer, 1997: 64 ( Gabrius ; characters; New Guinea).

- DISTRIBUTION: New Guinea.

praedictus Schillhammer, 1997: 57 ( Gabrius ; Type locality: Nepal: 328 Panchthar Distr. , Paniporua, 2300 m).

- DISTRIBUTION: Nepal.

primigenius Joy, 1913: 25 ( Gabrius ; Type locality: N.W. Spain, Pontevedra ).

— Scheerpeltz, 1933: 1367 ( Philonthus ; synonym of velox ).

— Smetana, 1957: 65 ( Gabrius ; synonym of velox ).

— Smetana, 1958: 251 ( Gabrius ; synonym of velox ).

— Smetana, 1960c: 331 ( Gabrius ; characters; Spain; North Africa; France).

— Smetana, 1967b: 159 ( Gabrius ; cited as promigenius; Spain).

— Coiffait, 1974: 60 ( Gabrius ; characters; North Africa; Iberian Peninsula; France).

— Pope, 1977: 30 ( Gabrius ; synonym of velox ).

— Smetana, 1995e: 627 ( Gabrius ; valid species).

— Outerelo, Gamarra, and Salgado, 1998: 129 ( Gabrius ; collected from cave; Spain).

- DISTRIBUTION: France, Spain, Morocco.

mauretanicus Peyerimhoff, 1937a: 266 ( Philonthus ; subgenus Gabrius ; Type locality: Causse du Mahmel, Aurès, vers 1800 m).

— Coiffait, 1951: 112, 115 ( Gabrius ; characters in key; France).

— Smetana, 1957: 65 ( Gabrius ; characters; North Africa; France).

— Smetana, 1960c: 331 ( Gabrius ; synonym of primigenius ).

— Smetana, 1967b: 159 ( Gabrius ; synonym of primigenius ).

— Coiffait, 1973a: 274 ( Gabrius ; Morocco).

— Coiffait, 1974: 60 ( Gabrius ; synonym of primigenius ).

propinquus Cameron, 1933d: 389 ( Philonthus ; subgenus Gabrius ; [preoccupied]; Type locality: Sumatra: Gunung Singgalang, 1000 m; [Note: See lectotype designation by Schillhammer, 1997]).

— Cameron, 1937: 6 ( Philonthus ; Java).

Schillhammer, 1997: 49 ( Gabrius ; lectotype designation; characters; Sumatra; Java; Bali) .

— Schillhammer, 2000 c: 132 ( Gabrius ; Java; Bali).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Indonesia.

pseudimitator Schillhammer, 1997: 46 ( Gabrius ; Type locality: Thailand: Chiang Mai,

Doi Suthep, 1550 m).

- DISTRIBUTION: Thailand.

pseudopennatus Cameron, 1945c: 716 ( Philonthus ; subgenus Gabrius ; Type locality:

South Africa: Simon’s Town).

— Tottenham, 1956: 305 ( Gabrius ; characters in key).

— Tottenham, 1956b: 207, 208 ( Gabrius ; characters).

- DISTRIBUTION: South Africa.

pubens Mulsant and Rey, 1878 , see: Gabronthus .

turneri Cameron, 1951c: 403 ( Philonthus ; subgenus Gabrius ; [preoccupied]; Type locality: Kenya Colony: Londiani).

— Schillhammer, 1997: 82 ( Gabrius ).

— Herman, 2001: 44 ( Gabrius ; synonym of pullatus ).

punctatellus Horn, 1884: 215 ( Philonthus ; [preoccupied]; Type locality: Ft. McLeod Brit. Amer.; [Note: See lectotype designation by Smetana, 1995e]. Originally cited from Hudson’s Bay region; Lake Tahoe ).

— Bernhauer and Schubert, 1914: 352 ( Philonthus ; catalog).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 692 ( Gabrius ; subspecies punctatellus ; lectotype designation: Ft. McLeod Brit. Amer.; characters; notes; Alberta; British Columbia; Saskatchewan; Yukon Territory; Colorado; Idaho; Montana; Oregon; Utah; Washington).

— Herman, 2001: 44 ( Gabrius ; although a junior primary homonym, prevailing use as valid maintained pending outcome of application to Commission under Article 23.9.5).

- DISTRIBUTION: Canada, USA.

ottawensis Casey, 1915: 438 ( Philonthus ; Type locality: Canada: Ottawa ; [Note: See lectotype designation by Smetana, 1995e]).

— Scheerpeltz, 1933: 1354 ( Philonthus ; catalog).

— Moore, 1965: 100 ( Gabrius ; cited as ottawaensis).

— Smetana, 1995e: 693, 695 ( Gabrius ; subspecies of punctatellus ; lectotype designation; characters; notes; Manitoba; Ontario; Quebec; Massachusetts; New Hampshire; New York; Wisconsin).

downiei Hatch, 1957: 203 ( Philonthus ; subgenus Gabrius ; Type locality: Washington: Pullman. Also cited from British Columbia and Idaho).

— Moore, 1965: 100 ( Gabrius ; cited as downei).

— Smetana, 1995e: 692 ( Gabrius ; synonym of punctatellus punctatellus ).

wawawai Hatch, 1957: 203 ( Philonthus ; subgenus Gabrius ; Type locality: Washington; Wawawai. Also cited from Oregon and British Columbia).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 692 ( Gabrius ; synonym of punctatellus punctatellus ).

miltiades Smetana, 1995e: 693 , 695 ( Gabrius ; subspecies of punctatellus ; Type locality: New Mexico: Lincoln N.F., 1 mi. SE Cloudcroft, 8750 feet. Also cited from Arizona, Nevada, Nebraska; labelled as intermediate between miltiades and punctatellus : Colorado and Utah).

pujoli Levasseur, 1967 , see: holisinus Fauvel, 1904 .

pumilus Mannerheim, 1830 , see: nigritulus Gravenhorst, 1802 .

pusillus Heer, 1839 , see: nigritulus Gravenhorst, 1802 .

puthzi Schillhammer, 1997: 35 ( Gabrius ; Type locality: Nepal [ Sindhupalchok District ]:

Prov. Bagmati, Malemchi, 2800 m).

- DISTRIBUTION: Nepal.

pygmaeus Stephens, 1833 , see: nigritulus Gravenhorst, 1802 .

pygmaeus Snellen, 1854 , see: nigritulus Gravenhorst, 1802 .

quadriceps Jarrige, 1978: 285 ( Gabrius ; Type locality: Andringitra Est, Anjavidilava, 2000 m). - DISTRIBUTION: Madagascar.

quadripunctatus Schillhammer, 1992: 64 ( Gabrius ; Type locality: China: Gansu, Meijishan ). - DISTRIBUTION: China.

rabigoides Jarrige, 1957: 110 ( Gabrius ; Type locality: Réunion: forêt de Bélouve).

— Lecoq, 1990: 192 ( Gabrius ; Mascarene Islands).

- DISTRIBUTION: Réunion.

ramifer Tottenham, 1956: 303 ( Gabrius ; Type locality: Ruanda: Shangugu , 1500 m).

- DISTRIBUTION: Rwanda. — Fagel, 1955a: 85 ( Gabrius ; Greece).

— Smetana, 1960c: 351 ( Gabrius ; characters from original description; species uncertain).

— Horion, 1965: 195 ( Gabrius ; Austria; Slovenia; Greece; Holland).

— Bordoni, 1973i: 329 ( Gabrius ; characters).

— Coiffait, 1974: 55 ( Gabrius ; characters; Europe; Asia Minor; Caucasus; Pyrenees).

— Smetana, 1977: 794 ( Gabrius ; Turkey).

— Tóth, 1984: 92 ( Gabrius ; characters; Hungary).

— Schillhammer, 1990: 91 ( Gabrius ; characters; notes).

— Smetana, 1993b: 49 ( Gabrius ; Czech Republic; Slovakia).

— Schillhammer, 1993: 326 ( Gabrius ; Austria).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; Italy).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; note; Austria).

— Tronquet, 1998: 11 ( Gabrius ; notes; France).

- DISTRIBUTION: Europe, Caucasus, Turkey.

spurius Smetana, 1954b: 178 ( Gabrius ; [Note: This name was validated by Smetana, 1954b; comparative characters were cited and an illustration of the aedeagus was published; the author stated the species was to be described later]; Type locality: Moravia : Prostějov [ Note : Localities taken from Smetana , 1956b: 175. Also cited from Anatolia, Bohemia, Czechoslovakia, Austria, Hungary and the Caucasus).

— Smetana, 1952b: 122 ( Gabrius ; [Note: Misidentified as subnigritulus according to Smetana, 1958: 250]; characters; Czechoslovakia).

— Smetana, 1957: 67, 69 ( Gabrius ; characters; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria; Caucasus; Turkey; Germany).

— Smetana, 1958: 250 ( Gabrius ; characters; ecology; Czechoslovakia).

— Smetana, 1960c: 323 ( Gabrius ; characters; Caucasus; Turkey; Czechoslovakia; Austria; Hungary; Transylvania; Bulgaria).

— Smetana, 1961a: 44 ( Gabrius ; Bulgaria).

— Lohse, 1964: 191 ( Gabrius ; characters; central Europe).

— Horion, 1965: 196 ( Gabrius ; Caucasus; Turkey; Bulgaria; Romania; Czech Republic; Slovakia; Austria).

— Smetana, 1967b: 159 ( Gabrius ; Spain).

— Coiffait, 1974: 55 ( Gabrius ; synonym of ravasinii ).

— Smetana, 1977: 794 ( Gabrius ; synonym of ravasinii ).

— Lucht, 1987: 106 ( Gabrius ; checklist; Czechoslovakia; Austria).

— Schillhammer, 1990: 91 ( Gabrius ; notes).

— Smetana, 1993b: 49 ( Gabrius ; synonym of ravasinii ).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; synonym of ravasinii ).

spurius Smetana, 1956b: 171 , 175 ( Gabrius ; [Note: Cited as a new species but the name was validated in Smetana, 1954b]; characters; Moravia. Also cited from Anatolia, Bohemia, Austria, Hungary, Transylvania, Bulgaria, and Caucasus).

nigrituloides Coiffait, 1963: 20 ( Gabrius ; [preoccupied]; Type locality: Forêt de Lente, Drôme).

— Coiffait, 1974: 55 ( Gabrius ; characters; France).

— Schillhammer, 1996a: 81 ( Gabrius ; synonym of ravasinii ).

riedeli Schillhammer, 1997: 70 ( Gabrius ; Type locality: Indonesia: N-Sumatra: Kutacane

Gg. Sinabung, 2000 m).

- DISTRIBUTION: Indonesia.

robustus Smetana, 1953a: 168 ( Gabrius ; Type locality: Macedonia: Kistac ).

— Smetana, 1957: 66 ( Gabrius ; characters; Macedonia).

— Smetana, 1960c: 349 ( Gabrius ; characters; Macedonia).

— Coiffait, 1974: 59 ( Gabrius ; characters; Macedonia).

— Schuh, Schillhammer, and Zettel, 1992: 220 ( Gabrius ; characters; notes; Austria).

— Assing, Frisch, Kahlen, et al., 1998: 138 ( Gabrius ; characters; Macedonia; Austria).

roubali Smetana, 1952a: 89 ( Gabrius ; Type locality: Rossia mer., Caucasus occ., Adler).

— Smetana, 1960c: 343 ( Gabrius ; characters; Caucasus).

— Coiffait, 1974: 59 ( Gabrius ; characters; Caucasus).

- DISTRIBUTION: Russia.

roubalioides Schillhammer, 1997: 15 ( Gabrius ; Type locality: O-Afghanistan: Do-Schak, Hindukusch, Khinjantal, 2500 m).

- DISTRIBUTION: Afghanistan.

rougemonti Schillhammer, 1997: 51 ( Gabrius ; Type locality: Bali: Penulisan ).

- DISTRIBUTION: Indonesia.

rubripennis Kiesenwetter, 1844 , see: lividipes Baudi, 1848 .

rufocinctus Schillhammer, 1997: 55 ( Gabrius ; Type locality: Nepal [Sindhupalchok District]: Prov. Bagmati, Malemchi , 2800 m).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: Nepal, India.

dubiosus Schillhammer, 1997: 56 ( Gabrius ; subspecies of rufocinctus ; Type locality: India: W. Bengal, Darjeeling dist., Tigerhill. Also cited from Nepal).

— Schillhammer, 2000 c: 133 ( Gabrius ; subspecies of rufocinctus ; Nepal).

rupicola Cameron, 1943: 131 ( Philonthus ; subgenus Gabrius ; Type locality: Darjeeling: Ghum district; [Note: See lectotype designation by Schillhammer, 1997]).

Schillhammer, 1997: 29 ( Gabrius ; lectotype designation; characters; India; Nepal) .

- DISTRIBUTION: India, Nepal.

sacerdotalis Joy, 1913: 26 ( Gabrius ; Type locality: Caucasus, Swanetia).

— Gridelli, 1920: 149 ( Gabrius ; characters; Caucasus).

— Scheerpeltz, 1933: 1359 ( Philonthus ; catalog).

— Smetana, 1960c: 339 ( Gabrius ; characters; Caucasus).

— Korge, 1971a: 7 ( Gabrius ; Turkey).

— Coiffait, 1974: 62 ( Gabrius ; characters; Caucasus).

- DISTRIBUTION: Turkey, Georgia.

sagittifer Schillhammer, 1997: 60 ( Gabrius ; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi Khola near Malemchi , 2100 m. Also cited from India).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: India, Nepal, China.

acutiphallus Schillhammer, 1997: 61 ( Gabrius ; subspecies of sagittifer ; Type locality: China: Yunnan, 100 km W. Baoshan, Gaoligongshan Nat. Res.).

sallustius Smetana, 1995e: 691 ( Gabrius ; Type locality: Oregon: Fly Cr. Valley , SE Umatilla County. Also cited from Washington).

— Hatch, 1957: 202 ( Philonthus ; subgenus Gabrius ; [Note: Misidentified as distans according to Smetana, 1995e: 689, 691]; characters; British Columbia; Washington).

- DISTRIBUTION: Canada, USA.

satanellus Bordoni, 1973 , see: doderoi Gridelli, 1920 .

schawalleri Coiffait, 1982 , see: furtivus Cameron, 1932 .

schawallerianus Schillhammer, 1997: 56 ( Gabrius ; Type locality: Nepal: 328 Panchthar Distr. , Paniporua, 2300 m).

— Schillhammer, 2000 c: 133 ( Gabrius ; Nepal).

- DISTRIBUTION: Nepal.

schillhammeri Shibata, 1996: 146 ( Gabrius ; replacement name for multipunctatus Shibata ). - DISTRIBUTION: Japan.

multipunctatus Shibata, 1991a: 88 ( Gabrius ; [preoccupied]; Type locality: Japan: Yamanashi Pref.: Akazawa Spa, Masuho ).

— Shibata, 1996: 146 ( Gabrius ; synonym of schillhammeri ).

schmidti Schillhammer, 2000 c: 123 View in CoL ( Gabrius View in CoL ; Type locality: Nepal Himalaya : Manaslu-Mts., Dudh Pokhari Lekh zw. Simia Kharka and Malamche Kharka, 33–3500 m).

- DISTRIBUTION: Nepal.

schoedli Schillhammer, 1997: 72 ( Gabrius ; Type locality: Indonesia: S-Sulawesi, E. Malino, Kanre Apia, 1400 m).

- DISTRIBUTION: Indonesia.

schwendingeri Schillhammer, 1997: 46 ( Gabrius ; Type locality: Thailand: Mae Hong Son - Doi Chang, 20 km E Pai).

- DISTRIBUTION: Thailand.

scrutandus Schillhammer, 1997: 35 ( Gabrius ; Type locality: Nepal: [Sankhua Sabha District]: Khandbari Distr., Induwa Khola Valley, 2850 m).

- DISTRIBUTION: Nepal.

seattlensis Hatch, 1957: 201 ( Philonthus ; subgenus Gabrius ; Type locality: Washington: Seattle. Also cited from British Columbia and Oregon).

— Moore, 1965: 100 ( Gabrius ).

— Smetana, 1995e: 669 ( Gabrius ; characters; notes; British Columbia; Oregon; Washington).

- DISTRIBUTION: Canada, USA.

semipunctatus Stephens, 1833: 252 ( Gabrius ; Type locality: Suffolk).

— Stephens, 1839: 401 ( Gabrius ; characters; England).

- DISTRIBUTION: Britain.

senegalensis Cameron, 1949a: 320 ( Gabrius ; Type locality: Guinée: Mt. Nimba , alt. 700–

900 m. Also cited from Ivory Coast).

- DISTRIBUTION: Guinea, Ivory Coast.

septempunctatus Cameron, 1930c: 207 ( Philonthus ; subgenus Gabrius ; Type locality: Kobe: Gomo).

— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).

— Cameron, 1949b: 468 ( Philonthus ; Formosa).

— Smetana, 1984: 122 ( Gabrius ; species unknown; Japan).

- DISTRIBUTION: Japan, Taiwan.

serpentis Schillhammer, 1997: 60 ( Gabrius ; Type locality: Nepal: [Sindhupalchok District]: Prov. Bagmati, Malemchi , 2800 m).

- DISTRIBUTION: Nepal.

sesengeanus Tottenham: [Note: Nomen nudum (Article 13.1.1); Tottenham, 1962: 143 cited this name in Gabrius , attributed it to himself, cited it from Uganda but without characters and without a reference to an earlier description].

sexualis Smetana, 1954: 42 ( Gabrius ; Type locality: Gallia mer., Alpes Maritimes, In. Vesubie. Also cited from Italy).

— Smetana, 1957: 63 ( Gabrius ; characters; France; Italy).

— Smetana, 1960c: 342 ( Gabrius ; characters; France; Italy).

Bordoni, 1973i: 327, 331 ( Gabrius ; characters; Italy) .

— Coiffait, 1974: 65 ( Gabrius ; characters; Alps; Italy; North Africa).

— Ciceroni and Zanetti, 1995: 29 ( Gabrius ; probable subspecies of coxalus ; Italy).

- DISTRIBUTION: Italy, France, “North Africa”.

sharpianus Cameron, 1930c: 207 ( Philonthus ; subgenus Gabrius ; Type locality: Settsu; Katsuogi: Nagasaki; Miyanoshita).

— Scheerpeltz, 1933: 1360 ( Philonthus ; catalog).

— Smetana, 1960c: 305 ( Gabrius ; characters; Japan).

— Smetana, 1984: 142 ( Gabrius ; characters; Japan).

- DISTRIBUTION: Japan, China.

conicus Bernhauer, 1939d: 99 ( Philonthus ; subgenus Gabrius ; Type locality: Japan: Tsushima. Nordwestl. China: Chinkiang).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Philonthus

Loc

Philonthus phaeopus

Herman, Taphylininae Iochini Aorothiini Thiini Latyprosopini Taphylinini Mblyopinina Nisolinina Yptiomina Hilonthina Lee H. 2001
2001
Loc

philomimus

Schillhammer 2000: 125
2000
Loc

schmidti

Schillhammer 2000: 123
2000
Loc

Gabrius

Stephens 1829
1829
Loc

Gabrius

Stephens 1829
1829
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