Sphex umtalicus Strand, 1916

Dörfel, Thorleif H. & Ohl, Michael, 2022, The wasp genus Sphex in Sub-Saharan Africa (Hymenoptera: Sphecidae), European Journal of Taxonomy 796 (1), pp. 1-170 : 72-80

publication ID

https://doi.org/10.5852/ejt.2022.796.1665

publication LSID

lsid:zoobank.org:pub:76C5C9C4-C6C1-4EDC-8FF8-9828A6EF2040

DOI

https://doi.org/10.5281/zenodo.6535796

persistent identifier

https://treatment.plazi.org/id/03C20928-FFD6-FFEC-4249-14EE384C09B1

treatment provided by

Felipe

scientific name

Sphex umtalicus Strand, 1916
status

 

Sphex umtalicus Strand, 1916

Figs 3–4 View Figs 1–6. 1–3 , 27 View Figs 25–32. 25–26 , 33 View Figs 33–40. 33–37 , 72, 77 View Figs 72–78. 72 (red)

Sphex umtalicus Strand, 1916: 105 , ♂.

Sphex nigripes var. pachyderma Strand, 1916: 106 , ♀. Syn. nov.

Chlorion haemorrhoidalis var. kobrowi Arnold, 1928: 367 , ♀ (holotype: ♀, South Africa, Gauteng, Johannesburg, TMP, not examined). Syn. nov.

Chlorion haemorrhoidalis var. basuto Arnold, 1947: 146 , sex not stated (holotype or syntypes: Lesotho, Berea District, Mamathes, SAM, not examined). Syn. nov.

Differential diagnosis

In most cases, specimens of S. umtalicus are easily identifiable within the species group through their black clypeus, uniformly black legs and fuscous wings with cyan-violet iridescence ( Fig. 72 View Figs 72–78. 72 ). However, we observed intraspecific variations in this species as well as in S. haemorrhoidalis that can cause confusion between the two of them. Females of S. haemorrhoidalis normally have the clypeus ferruginous ( Fig. 34 View Figs 33–40. 33–37 ), though it can also be black to a varying extent, sometimes almost completely. While the tibiae and part of the femora and tarsi are always ferruginous in female S. haemorrhoidalis ( Fig. 69 View Figs 63–71. 63, 67 ) and S. victoria sp. nov. ( Fig. 73 View Figs 72–78. 72 ), a southern African population of S. umtalicus (previously described as var. basuto / kobrowi ) also has these areas ferruginous, as well as the lower part of the female clypeus. The best way to differentiate between these females is the color of the appressed clypeal and paraocular setae, which is always dark brown in S. umtalicus and can have a slight silvery shimmer on the paraocular area ( Fig. 33 View Figs 33–40. 33–37 ). The appressed clypeal setae of female S. haemorrhoidalis are inconspicuous, but the appressed lateroclypeal and paraocular setae have a ferruginous-golden shimmer ( Fig. 34 View Figs 33–40. 33–37 ).

Males of the southern African population of S. umtalicus are characterized by having the wings mostly hyaline and all tibiae ferruginous, whereas those of the other species in the group have only the hindtibia ferruginous. Like all males of the species, they have silvery appressed clypeal setae and uniformly black erect mesosomal setae.

Material examined

Holotype

ZIMBABWE – Manicaland • ♂; Umtali [now Mutare] ; A. Bodong leg.; DEI.

Other material

AFRICA • 1 ♀; “Afr. or. All. de”; MNHN 1 ♀; “ East Africa ”; IRSN 1 ♀; “ Eersteleng ”; BMNH 3 ♂♂; “ Lake Kivu ”; R. Kandt leg.; ZMB 1 ♂; “ Tanganyika ”; H.A. Lindeman leg.; BMNH 1 ♀; German East Africa ; 1912; MNHN 1 ♀; same collection data as for preceding; MNHN .

ANGOLA – Cuanza Sul Province • 1 ♀; Congulu ; [11°17ʹ03ʺ S, 15°45ʹ08ʺ E]; Apr. 1934; K. Jordan leg.; BMNH GoogleMaps .

DEMOCRATIC REPUBLIC OF THE CONGO – Haut-Katanga • 1 ♀; Sakania ; [12°45ʹ S, 28°33ʹ36ʺ E]; 15 Mar. 1938; IRSN GoogleMaps 1 ♀; Sakania ( marais Lukabungu ); [12°45ʹ S, 28°34ʹ E]; 15 Mar. 1938; H.J. Brédo leg.; MRAC GoogleMaps 1 ♀; Tera ; [12°08ʹ S, 28°34ʹ E]; Feb. 1948; R. Mouchamps leg.; MRAC GoogleMaps . – Haut-Lomami • 1 ♂; “ Upemba National Park, Kankunda ( rive dr. Lupiala )”; [8°30ʹ S, 26°30ʹ E]; alt. 1300 m; 13–27 Nov. 1947; G.F. de Witte leg.; IRSN GoogleMaps 1 ♂; Lemfu ; [5°17ʹ56ʺ S, 15°13ʹ21ʺ E]; May 1945; P.L. De Beir leg.; MRAC GoogleMaps 1 ♀; Thysville [ now Mbanza-Ngungu ]; 5°30ʹ S, 15°00ʹ E; 2 Jun. 1915; Lang and Chapin leg.; AMNH GoogleMaps . – Kongo Central Province • 2 ♂♂; Mayumbe ; Nov. 1938; IRSN .

ETHIOPIA – Amhara Region • 1 ♂; Jowaha , N of Debra Sina; 7 Aug. 1946; K.M. Guichard leg.; BMNH .

KENYA – Kilifi County • 1 ♂, 2 ♀♀; Rabai ; [3°55ʹ54ʺ N, 39°34ʹ13ʺ E]; Apr. 1928; V.G.L. van Someren leg.; BMNH GoogleMaps . – Kwale County • 3 ♂♂; Funzi Island, near Funzi workshop; 4.57749° S, 39.43825° E; 4–10 Jul. 2012; ICIPE / NMK leg.; ICIPE GoogleMaps 1 ♀; Funzi Island, near Mliani; 4.53577° S, 39.46042° E; 6 Jul. 2012; ICIPE / NMK leg.; ICIPE GoogleMaps 1 ♀; Funzi Island , near swimming pool; 4.58463° S, 39.43916° E; 9 Jul. 2012; ICIPE/NMK leg.; ICIPE GoogleMaps 1 ♂; Shimba Hills ; 4°12.3ʹ S, 39°25ʹ E; 15 Dec. 2003; W.J. Pulawski leg.; CAS GoogleMaps 1 ♂; Shimba Hills (nr. Kwale); [4°16ʹ31ʺ S, 39°23ʹ45ʺ E]; 13 May 1979; R. Murphy leg.; BMNH GoogleMaps 1 ♂; Tiwi Beaches , ca 17 km S of Mombasa; 4°14.4ʹ S, 39°36.2ʹ E; 24–26 Jul. 1999; W.J. Pulawski and J.S. Schweikert leg.; CAS GoogleMaps . – Makueni County • 1 ♂; Makueni, Ukamba ; [1°48ʹ S, 37°37ʹ E]; May 1947; V.G.L. van Someren leg.; BMNH GoogleMaps . – Nairobi County • 1 ♀; near Nairobi, Vte. de Pongins et Cte de Lambertye ; [1°17ʹ S, 36°49ʹ E]; MNHN GoogleMaps . – Taita-Taveta County • 4 ♀♀; Kasigau ; [3°49ʹ S, 38°40ʹ E]; Nov. 1938; BMNH GoogleMaps 1 ♀; near Voi, Taita Hills ; [3°23ʹ26ʺ S, 38°34ʹ37ʺ E]; 16–18 Nov. 1921; H.E. Box leg.; BMNH GoogleMaps 1 ♀; Voi ; [3°23ʹ26ʺ S, 38°34ʹ37ʺ E]; 23 Nov. 1997; M. Snižek leg.; OÖLM GoogleMaps 3 ♀♀; Wundanyi ; [3°23ʹ54ʺ S, 38°21ʹ37ʺ E]; 19–21 Nov. 1996; Mi. Halada leg.; OÖLM GoogleMaps 10 ♂♂, 1 ♀; same locality as for preceding; 5–10 Apr. 1997; Ma. Halada leg.; OÖLM GoogleMaps 3 ♂♂, 1 ♀; same collection data as for preceding but 18–22 Mar. 1997; OÖLM GoogleMaps .

LESOTHO – Berea District • 1 ♀; Mamathes ; [29°08ʹ S, 27°51ʹ E]; 28 Dec. 1946; C.F. Jacot-Guillarmod leg.; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 1 Jan. 1947; BMNH GoogleMaps .

MALAWI • 1 ♀; NW shore of Lake Nyasa, between Florence Bay and Karonga; 30 Jun.–6 Jul. 1910; S.A. Neave leg.; BMNH. – Central Region • 1 ♀; 100 km S of Kasungu; [13°57ʹ37.2ʺ S, 33°28ʹ16.5ʺ E]; 29 Dec. 2001; J. Halada leg.; OÖLM GoogleMaps 1 ♀; Dedza, 85 km SE of Lilongwe; [14°20ʹ S, 34°20ʹ E]; 17–19 Dec. 2001; J. Halada leg.; OÖLM. – GoogleMaps Northern Region • 1 ♀; Karonga; [9°56ʹ S, 33°56ʹ E]; 7–11 Jul. 1910; S.A. Neave leg.; BMNH. – GoogleMaps Southern Region • 1 ♀; Mlanje ; [16°01ʹ53ʺ S, 35°30ʹ E]; 23 Jan. 1912; S.A. Neave leg.; BMNH GoogleMaps 2 ♂♂; same collection data as for preceding but 4 Dec. 1912; BMNH GoogleMaps 2 ♂♂; same collection data as for preceding but 6 Dec. 1912; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 11 Dec. 1912; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 12 Dec. 1912; BMNH GoogleMaps 3 ♂♂, 1 ♀; same collection data as for preceding but 14 Dec. 1912; BMNH GoogleMaps 1 ♂, 1 ♀; same collection data as for preceding but 23 Dec. 1912; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 26 Dec. 1912; BMNH GoogleMaps 3 ♂♂, 2 ♀♀; same collection data as for preceding but 6 Jan. 1913; BMNH GoogleMaps 6 ♂♂, 1 ♀; same collection data as for preceding but 8 Jan. 1913; BMNH GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 13 Jan. 1913; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 14 Jan. 1913; BMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but 15 Jan. 1913; BMNH GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 17 Jan. 1913; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 17 Jan. 1913; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 21 Jan. 1913; BMNH GoogleMaps 3 ♂♂; same collection data as for preceding but 22 Jan. 1913; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 24 Jan. 1913; BMNH GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 27 Jan. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 29 Jan. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 1 Feb. 1913; BMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but 4 Feb. 1913; BMNH GoogleMaps 1 ♂, 1 ♀; same collection data as for preceding but 6 Feb. 1913; BMNH GoogleMaps 1 ♂, 1 ♀; same collection data as for preceding but 8 Feb. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 11 Feb. 1913; BMNH GoogleMaps 1 ♂, 1 ♀; same collection data as for preceding but 14 Feb. 1913; BMNH GoogleMaps 5 ♀♀; same collection data as for preceding but 17 Feb. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 24 Feb. 1913; BMNH GoogleMaps 2 ♀♀; same collection data as for preceding but 27 Feb. 1913; BMNH GoogleMaps 2 ♂♂, 3 ♀♀; same collection data as for preceding but 3 Mar. 1913; BMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but 5 Mar. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 20 Mar. 1913; BMNH GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 26 Mar. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 17 Apr. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 10 May 1913; BMNH GoogleMaps 2 ♀♀; same collection data as for preceding; BMNH GoogleMaps 3 ♂♂, 2 ♀♀; same collection data as for preceding but 6 Dec. 1913; BMNH GoogleMaps 1 ♂, 5 ♀♀; same collection data as for preceding but 21 Dec. 1913; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 29 Dec. 1913; BMNH GoogleMaps 3 ♀♀; same collection data as for preceding but 20 Jan. 1914; BMNH GoogleMaps 2 ♂♂, 6 ♀♀; same collection data as for preceding but 27 Jan. 1914; BMNH GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 10 Feb. 1914; BMNH GoogleMaps 2 ♂♂; same collection data as for preceding but 13 Feb. 1914; BMNH GoogleMaps 6 ♀♀; same collection data as for preceding but 16 Feb. 1914; BMNH GoogleMaps 6 ♂♂, 1 ♀; side of Mt Mlanje; [16°01ʹ53ʺ S, 35°30ʹ E]; 3 Dec. 1913; S.A. Neave leg.; BMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but 11 Dec. 1913; BMNH GoogleMaps 1 ♀; side of Mt Mulanje; [16°01ʹ53ʺ S, 35°30ʹ E]; alt. 3000–4000 ft; 2 Dec. 1913; S.A. Neave leg.; MNHN GoogleMaps 5 ♀♀; SW of Lake Chilwa; [15°24ʹ01ʺ S, 35°34ʹ47ʺ E]; 9 Jan. 1914; S.A. Neave leg.; BMNH GoogleMaps 1 ♀; Zomba; [15°23ʹ09ʺ S, 35°19ʹ07ʺ E]; Nov. 1913; H.S. Stannus leg.; BMNH GoogleMaps 1 ♀; same collection data as for preceding but Dec. 1913; BMNH GoogleMaps 2 ♀♀; same collection data as for preceding but Jan. 1914; BMNH GoogleMaps 1 ♀; same locality as for preceding but Dec. 1896; BMNH GoogleMaps 1 ♀; Zomba Mt ; [15°17ʹ S, 35°17ʹ E]; alt. 5500 ft; 18 Feb. 1910; S.A. Neave leg.; BMNH GoogleMaps .

MOZAMBIQUE • 1 ♀; valley of Kola River, near E Mt Chiperone; 3Apr. 1913; S.A. Neave leg.; BMNH 1 ♀; same collection data as for preceding but 6 Apr. 1913; BMNH 2 ♀♀; same collection data as for preceding but 7 Apr. 1913; BMNH. – Manica Province • 1 ♀; 65 km W of Chimoio; [18°57ʹ51ʺ S, 32°50ʹ10.1ʺ E]; 10–11 Feb. 2003; J. Halada leg.; OÖLM GoogleMaps 1 ♀; Amatongas Forest ; [19°11ʹ S, 33°45ʹ E]; Feb. 1917; NRS GoogleMaps 1 ♀; near Andrada, Valee du Revoué ; [18°53ʹ42.6ʺ S, 32°51ʹ16.7ʺ E]; 1905; G. Vasse leg.; MNHN GoogleMaps 1 ♂; Vila Pery (now Chimoio ); [19°06ʹ59ʺ S, 33°29ʹ E]; 1928; P. Lesne leg.; MNHN. – GoogleMaps Tete Province • 1 ♂; 65 km S of Vlóngné; 15°13ʹ S, 34°19ʹ E; 8 Feb. 2005; J. Halada leg.; OÖLM. – GoogleMaps Zambezia Province • 2 ♂♂; Ruo Valley ; alt. 2000 ft; 1 Apr. 1913; S.A. Neave leg.; BMNH GoogleMaps 1 ♀; same collection data as for preceding; MNHN GoogleMaps 1 ♂; same collection data as for preceding but 14 Dec. 1913; BMNH GoogleMaps .

NAMIBIA – Kunene Region • 1 ♀; 38 km NW of Outjo; [19°49ʹ53ʺ S, 15°55ʹ33ʺ E]; 6 Mar. 1990; W.J. Pulawski leg.; CAS GoogleMaps . – Otjozondjupa Region • 1 ♀; Otavi ; [19°39ʹ S, 17°20ʹ E]; 2 Feb. 1993; M. Schwarz leg.; OÖLM GoogleMaps .

SOUTH AFRICA • 1 ♀; Pienaars River ; 1898; R. v. Jutrzencka leg.; TMP. – Eastern Cape • 1 ♀; Gonubie ; 32°55ʹ49.40ʺ S, 28°01ʹ5.41ʺ E; 21 Jan. 2013; C.P. Fielding leg.; ZMB GoogleMaps 1 ♂; Port St John, Pondoland ; [31°37ʹ22ʺ S, 29°32ʹ41ʺ E]; Nov. 1923; R.E. Turner leg.; BMNH GoogleMaps 1 ♂; same collection data as for preceding but Jan. 1924; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 6–25 Feb. 1924; BMNH. – GoogleMaps Gauteng • 1 ♀; Boukenhoutkloof ; 18 Dec. 1910; J.V. Niekerk leg.; TMP GoogleMaps 2 ♂♂; Pretoria ; [25°44ʹ46ʺ S, 28°11ʹ17ʺ E]; 21 Dec. 1926; W. Lingnau leg.; DEI GoogleMaps 1 ♀; same locality as for preceding; O. Ulbrich leg.; ZMB GoogleMaps 1 ♀; same locality as for preceding; W. Lingnau leg.; DEI GoogleMaps 2 ♂♂, 7 ♀♀; same locality as for preceding; W.L. Distant leg.; BMNH GoogleMaps 1 ♀; Pretoria City ; [25°43ʹ32ʺ S, 28°14ʹ38ʺ E]; 12 Feb. 2000; J. Halada leg.; OÖLM. – GoogleMaps KwaZulu-Natal • 1 ♂; MNHN GoogleMaps 1 ♀; 10 miles W of Durban; [29°50ʹ08.5ʺ S, 30°51ʹ2.4ʺ E]; 4 Dec. 1966; J.G. Rozen and D.J. Brothers leg.; AMNH GoogleMaps 1 ♂; 20 km NE of Ixopo; 30°00ʹ S, 30°12ʹ E; 15 Jan. 2010; J. Halada leg.; OÖLM GoogleMaps 2 ♂♂, 2 ♀♀; 20 km S of Bergville; [28°54ʹ49.2ʺ S, 29°21ʹ10.7ʺ E]; 5 Feb. 2001; M. Snižek leg.; OÖLM GoogleMaps 4 ♂♂; 20 km S of Newcastle; [27°56ʹ09.7ʺ S, 29°58ʹ12ʺ E]; 28 Nov. 2002; Ma. Halada leg.; OÖLM GoogleMaps 6 ♂♂, 2 ♀♀; 20 km W of Port Shepstone; [30°46ʹ50.9ʺ S, 30°12ʹ38ʺ E]; 2 Feb. 2000; J. Halada leg.; OÖLM GoogleMaps 5 ♂♂; 20 km W of Vryheid; [27°46ʹ28.8ʺ S, 30°34ʹ19.7ʺ E]; 29 Nov. 2002; Ma. Halada leg.; OÖLM GoogleMaps 1 ♂, 2 ♀♀; 35 km N of Port Edward; 30°45ʹ S, 30°09ʹ E; 28 Dec. 2009; J. Halada leg.; OÖLM GoogleMaps 1 ♀; Drakensberge, Royal Natal National Park ; 28°43ʹ S, 28°53ʹ E; 9 Feb. 1994; F. Koch leg.; ZMB GoogleMaps 1 ♂; Durban ; [29°53ʹ S, 31°03ʹ E]; J. Bequaert leg.; MRAC GoogleMaps 1 ♂; same collection data as for preceding; AMNH GoogleMaps 1 ♀; same collection data as for preceding; BMNH GoogleMaps 1 ♀; Durban, Congella ; [29°53ʹ S, 31°03ʹ E]; 2 May 1908; G.F. Leigh leg.; TMP GoogleMaps 1 ♀; Estcourt ; [29°00ʹ S, 29°53ʹ E]; 5 Mar. 1963; H.N. Empey leg.; RMNH GoogleMaps 1 ♀; Hluhluwe Game Reserve ; 28°02ʹ S, 32°05ʹ E; 10 Feb. 1995; F. Koch leg.; ZMB GoogleMaps 1 ♀; same collection data as for preceding but 5 Dec. 1995; ZMB GoogleMaps 2 ♂♂, 9 ♀♀; Howick ; [29°28ʹ40ʺ S, 30°13ʹ50ʺ E]; 1904; J.P. Cregoe leg.; BMNH GoogleMaps 1 ♂, 1 ♀; Ingogo , N of Newcastle; [27°34ʹ40.8ʺ S, 29°54ʹ43.2ʺ E]; 28 Nov. 2002; Ma. Halada leg.; OÖLM GoogleMaps 1 ♂; Ithala Game Reserve ; 27°30ʹ S, 31°20ʹ E; 28–30 Jan. 1995; F. Koch leg.; THD-002-ZMB; GenBank CO1 gene: MW538571 View Materials ; GenBank EF-1α gene: MW558237 View Materials ; GenBank LWR gene: MW582277 View Materials ; ZMB GoogleMaps 1 ♂; same collection data as for preceding; ZMB GoogleMaps 1 ♂, 2 ♀♀; same collection data as for preceding but 19 Feb. 1997; ZMB GoogleMaps 2 ♂♂, 4 ♀♀; Ithala Game Reserve, Louwsburg ; [27°32ʹ48.14ʺ S, 31°18ʹ48.71ʺ E]; 10–23 Dec. 1993; F. Koch leg.; ZMB GoogleMaps 2 ♀♀; Ithala Game Reserve , sports ground ; 27°30ʹ S, 31°20ʹ E; 5 Apr. 2001; F. Koch leg.; ZMB GoogleMaps 1 ♀; Lake St Lucia, False Bay ; 27°58ʹ S, 32°23ʹ E; 5–15 Feb. 1992; A.J. Weaving leg.; AMG GoogleMaps 1 ♂, 4 ♀♀; Malvern ; [29°53ʹ06ʺ S, 30°55ʹ30ʺ E]; 1904; J.P. Cregoe leg.; BMNH GoogleMaps 1 ♂; Maputaland , SW of Manguzi; [27°00ʹ48.5ʺ S, 32°43ʹ24.1ʺ E]; 29 Jan. 2003; M. Snižek leg.; OÖLM GoogleMaps 2 ♂♂; Mkuze Game Reserve ; 27°40ʹ S, 32°19ʹ E; 22 Nov. 1999; F. Koch leg.; ZMB GoogleMaps 1 ♀; Ndumo Game Reserve ; 26°55ʹ S, 32°19ʹ E; 19 Nov. 1999; F. Koch leg.; ZMB GoogleMaps 1 ♀; Pinetown ; [29°49ʹ14ʺ S, 30°53ʹ12ʺ E]; 18 Mar. 1909; G.F. Leigh leg.; OUMNH GoogleMaps 1 ♂; Pongola ; [27°22ʹ41ʺ S, 31°37ʹ08ʺ E]; 10 May 1951; H.W. Bedford leg.; BMNH GoogleMaps 3 ♂♂, 3 ♀♀; St Lucia Park ( Charter’s Creek ); 28°12ʹ S, 32°25ʹ E; 7 Feb. 1995; F. Koch leg.; ZMB GoogleMaps 1 ♀; Stanger [now KwaDukuza]; [29°20ʹ S, 31°17ʹ30ʺ E]; 14 Mar. 1963; H.N. Empey leg.; RMNH GoogleMaps 8 ♂♂; Utrecht ; [27°39ʹ36ʺ S, 30°19ʹ48ʺ E]; 28 Nov. 2002; M. Snižek leg.; OÖLM GoogleMaps 1 ♀; Weenen ; [28°50ʹ57ʺ S, 30°04ʹ38ʺ E]; Jan.–Mar. 1927; H.P. Thomasset leg.; BMNH. – GoogleMaps Limpopo • 1 ♂, 2 ♀♀; 35 km W of Hoedspruit; 24°23ʹ S, 30°40ʹ E; 19 Dec. 2009; J. Halada leg.; OÖLM GoogleMaps 1 ♂, 1 ♀; Lekgalameetse Nature Reserve ; 24°12ʹ S, 30°20ʹ E; 25–31 Mar. 2001; F. Koch leg.; ZMB GoogleMaps 1 ♀; Moorddrift ; [24°16ʹ5.8ʺ S, 28°57ʹ52.3ʺ E]; Oct. 1909; C.J. Swierstra leg.; TMP GoogleMaps 4 ♂♂, 3 ♀♀; N of Nylstroom [now Modimolle ]; 24°40ʹ S, 28°30ʹ E; 14 Jan. 2008; M. Halada leg.; OÖLM GoogleMaps 1 ♂; Thabazimbi ; [24°36ʹ S, 27°24ʹ E]; 21 Dec. 2008; M. Snižek leg.; OÖLM. – GoogleMaps Mpumalanga • 1 ♂, 1 ♀; 20 km NW of Nelspruit [now Mbombela ]; [25°20ʹ09ʺ S, 30°49ʹ35.5ʺ E]; 26 Nov. 2003; J. Halada leg.; OÖLM GoogleMaps 1 ♀; 20 km SE of Nelspruit [now Mbombela ]; [25°36ʹ41.7ʺ S, 31°05ʹ35.3ʺ E]; 5–6 Jan. 2004; J. Halada leg.; OÖLM GoogleMaps 2 ♀♀; Barberton ; [25°47ʹ18ʺ S, 31°03ʹ11ʺ E]; P. Rendall leg.; BMNH GoogleMaps 1 ♀; Nwaswitshaka Camp, Kruger National Park ; [24°59ʹ45ʺ S, 31°35ʹ31ʺ E]; 21 Dec. 1994; C.L. Bellamy leg.; AMNH GoogleMaps 1 ♀; Waterval River ; 8 Jan. 1903; J. Breyer leg.; TMP. – North West • 1 ♂, 2 ♀♀; Rustenburg ; [25°40ʹ03ʺ S, 27°14ʹ31ʺ E]; BMNH GoogleMaps 1 ♂; Sun City ; 32°37ʹ S, 19°07ʹ E; 24 Jan. 2006; A.R. Davidson leg.; AMNH GoogleMaps .

TANZANIA • 1 ♀; Tabora-Kigoma; Stamper leg.; MRAC. Dar es Salaam Region • 1 ♀; Dar es Salaam; [6°48ʹ S, 39°17ʹ E]; 1896; H. Mayer leg.; IRSN. GoogleMaps Katavi Region • 1 ♂, 1 ♀; 30 km NNW of Mpanda; 5°54ʹ S, 30°56ʹ E; 30 Dec. 2006; M. Kadlecová leg.; OÖLM. GoogleMaps Kigoma Region • 1 ♂; 25 km SSE of Kasulu; 4°44ʹ S, 30°11ʹ E; 28 Dec. 2006; J. Halada leg.; OÖLM. GoogleMaps Kilimanjaro Region • 1 ♀; Kilimanjaro, Obstgarten-Steppe ; [3°05ʹ5.9ʺ S, 37°20ʹ38.5ʺ E]; 24 Mar. 1905; Y. Sjöstedt leg.; BMNH. GoogleMaps Lindi Region • 3 ♀♀; Lindi; [9°59ʹ49ʺ S, 39°42ʹ59ʺ E]; F. Fülleborn leg.; ZMB. GoogleMaps Mbeya Region • 1 ♀; Langenburg [now Tukuyu ]; [9°15ʹ S, 33°39ʹ E]; end of Feb.–beginning of Mar. 1898; F. Fülleborn leg.; ZMB GoogleMaps 1 ♂; same collection data as for preceding but 3 Sep–3 Oct. 1898; ZMB • 1 ♂; Manow ; [9°15ʹ S, 33°48ʹ E]; MNHN GoogleMaps 1 ♂; Mbeya city; [8°54ʹ S, 33°27ʹ E]; 5 Jan. 2007; J. Halada leg.; OÖLM. GoogleMaps Mjini Magharibi Region • 2 ♀♀; Zanzibar; [6°08ʹ S, 39°20ʹ E]; 1872–1874; Hildebrandt leg.; ZMB GoogleMaps 1 ♀; same locality as for preceding; Dec. 1932; A. Seyrig leg.; MNHN GoogleMaps . – Rukwa Region • 2 ♀♀; Msamwia, S of Ufipa, S Tanganyika; [8°25ʹ S, 31°50ʹ E]; 10–12 Feb. 1909; Fromm leg.; ZMB. GoogleMaps Ruvuma Region • 1 ♀, lectotype of Sphex nigripes var. pachyderma Strand, 1916 (designated by Oehlke & Wudowenz 1973: 424); Kigonsera, Konow; DEI. Singida Region • 1 ♂; 20 mi. SW of Itigi; [5°53ʹ32ʺ S, 34°16ʹ02ʺ E]; 28 Jan. 1970; M.E. Irwin and E.S. Ross leg.; CAS. GoogleMaps Songwe Region • 1 ♂; 70 km NW of Tunduma; 8°58ʹ S, 32°25ʹ E; 2 Jan. 2007; J. Halada leg.; OÖLM. GoogleMaps Tanga Region • 1 ♀; “ Mnoa ”; F. Fischer leg.; ZMB • 1 ♀; Tanga; [5°04ʹ08ʺ S, 39°05ʹ55ʺ E]; 16 Dec. 1917; W.A. Lamborn leg.; OUMNH. GoogleMaps Unguja North Region • 1 ♂; Nungwi , Zanzibar; [5°43ʹ33.6ʺ S, 39°17ʹ45.6ʺ E]; 18 Jun. 1985; G.G.M. Schulten leg.; RMNH. GoogleMaps

UGANDA – Eastern Region • 1 ♂; between Jinja and Busia or Mbwago’s E Busora (Some Forest); [0°15ʹ44ʺ N, 33°35ʹ17ʺ E]; 28 Jul.–1 Aug. 1911; S.A. Neave leg.; BMNH GoogleMaps 1 ♂; Mt Kokanjero , SW of Elgon; [1°07ʹ48ʺ N, 34°33ʹ E]; 7–9 Aug. 1911; S.A. Neave leg.; BMNH GoogleMaps .

ZAMBIA • 1 ♀; “ Mpika – Fort Jameson ”; Jan. 1908; S.A. Neave leg.; MRAC 1 ♀; Chingola–Solwezi Road ; 2 Dec. 2004; M. Snižek leg.; OÖLM. – Central Province • 2 ♂♂; 100 km NE of Kapiri Mposhi; [13°36ʹ29.5ʺ S, 29°31ʹ49.3ʺ E]; 4–7 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps 1 ♂; 60 km NW of Kapiri Mposhi; [13°32ʹ08.4ʺ S, 28°18ʹ42.6ʺ E]; 8 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps 1 ♂; 70 km NE of Kapiri Mposhi; [13°41ʹ S, 29°18ʹ E]; 21 Nov. 2005; M. Snižek leg.; OÖLM GoogleMaps 1 ♀; Broken Hill [now Kabwe]; [14°26ʹ48ʺ S, 28°26ʹ47ʺ E]; 27 Feb. 1910; Silverlock leg.; BMNH. – GoogleMaps Copperbelt Province • 1 ♀; 45 km SE of Kitwe; [13°08ʹ36.7ʺ S, 28°32ʹ54.4ʺ E]; 12–15 Jan. 2003; J. Halada leg.; OÖLM GoogleMaps 3 ♂♂, 1 ♀; 50 km W of Chingola; [12°25ʹ06.3ʺ S, 27°23ʹ50.9ʺ E]; 9–11 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps 2 ♀♀; same collection data as for preceding but 1–2 Jan. 2003; OÖLM GoogleMaps 1 ♂; 80 km W of Chingola; [12°30ʹ33.2ʺ S, 27°06ʹ11.2ʺ E]; 12 Dec. 2002; F. Kantner leg.; OÖLM GoogleMaps 1 ♂; Tanga Farm , 35 km W of Chingola; [12°32ʹ27.7ʺ S, 27°31ʹ59.4ʺ E]; 6 Nov. 2005; M. Snižek leg.; OÖLM. – GoogleMaps Eastern Province • 1 ♂; “ Mid. Luangwa Valley ”; alt. 1700 ft; 26 Feb. 1908; S.A. Neave leg.; OUMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but 27 Feb. 1908; OUMNH GoogleMaps 4 ♀♀; “ On road Ft. Jameson to Lundazi ”; 7–14 Jun. 1910; S.A. Neave leg.; BMNH 1 ♀; “ Up. Luangwa Valley ”; alt. 1800–2000 ft; 30 Mar. 1908; S.A. Neave leg.; OUMNH 2 ♀♀; same collection data as for preceding but 31 Mar. 1908; OUMNH 1 ♂; same collection data as for preceding but 1 Apr. 1908; OUMNH 1 ♀; 6 km E of Sinda; 14°12ʹ S, 31°49ʹ E; 5 Mar. 1998; W.J. Pulawski leg.; CAS GoogleMaps 1 ♀; Petauke ; [14°14ʹ28ʺ S, 31°19ʹ11ʺ E]; 2 Feb. 1905; S.A. Neave leg.; OUMNH. – GoogleMaps Lusaka Province • 3 ♀♀; 25 km E of Lusaka; 15°21ʹ S, 28°30ʹ E; 3 Mar. 1995; W.J. Pulawski leg.; CAS. – GoogleMaps Muchinga Province • 2 ♀♀; “ Mid. Chambezi Valley, Chinsali distr. ”; alt. 4000 ft; 25 Apr. 1908; S.A. Neave leg.; OUMNH GoogleMaps 1 ♀; same collection data as for preceding but 27 Apr. 1908; OUMNH GoogleMaps 1 ♀; same collection data as for preceding but 29 Apr. 1908; OUMNH GoogleMaps 1 ♀; Chinsali and neighbourhood ; [10°32ʹ29ʺ S, 32°04ʹ53ʺ E]; alt. 4300 ft; 9 Apr. 1908; S.A. Neave leg.; OUMNH. – GoogleMaps Northern Province • 1 ♀; “ L. Chambezi V. ; Kasama distr. ”; alt. 3900 ft; 10 May 1908; S.A. Neave leg.; OUMNH GoogleMaps 1 ♀; 5 mi. S of Senga Hill; [9°26ʹ44ʺ S, 31°13ʹ21ʺ E]; 1 Feb. 1970; M.E. Irwin and E.S. Ross leg.; CAS GoogleMaps 1 ♂; Abercorn [now Mbala]; [8°50ʹ S, 31°28ʹ E]; 23 Dec. 1950; F.O. Albrecht leg.; BMNH GoogleMaps 1 ♂; same collection data as for preceding but 10 Jan. 1951; BMNH. – GoogleMaps Southern Province • 1 ♂; 10 km SW of Choma; [16°51ʹ57ʺ S, 26°54ʹ06.3ʺ E]; 18 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps 1 ♀; 100 km NE of Livingstone; [17°06ʹ21.1ʺ S, 26°24ʹ21ʺ E]; 26 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps 1 ♀; 30 km SE of Choma; [16°54ʹ09.4ʺ S, 27°15ʹ54.8ʺ E]; 27 Dec. 2002; J. Halada leg.; OÖLM GoogleMaps .

ZIMBABWE • 1 ♂; “ Murntree ”; 21 Jan. 1912; G. Arnold leg.; BMNH 1 ♀; Victoria Falls ; Feb. 1935; NRS. – Harare Province • 1 ♀; near Harare; [17°49ʹ45ʺ S, 31°03ʹ08ʺ E]; 20 Jan. 1998; Ma. Halada leg.; OÖLM GoogleMaps 1 ♀; Salisbury [now Harare]; [17°49ʹ45ʺ S, 31°03ʹ08ʺ E]; Apr. 1899; G.A.K. Marshall leg.; BMNH GoogleMaps 2 ♂♂, 1 ♀; same collection data as for preceding but Jan. 1900; BMNH GoogleMaps 1 ♀; same collection data as for preceding but 13 Dec. 1902; OUMNH GoogleMaps 3 ♀♀; same locality as for preceding; A. Watsham leg.; BMNH. – GoogleMaps Mashonaland Central • 1 ♀; 10 mi. E of Pakasa [ Bakasa ]; [16°25ʹ39ʺ S, 30°50ʹ32ʺ E]; 28 Jan. 1911; Silverlock leg.; BMNH GoogleMaps 5 ♂♂; Mavhuradonha Saf. , 15 km SE of Muzarabani; [16°33ʹ33ʺ S, 31°14ʹ22.8ʺ E]; 17 Dec. 1998; J. Halada leg.; OÖLM. – GoogleMaps Mashonaland East • 2 ♀♀; Chishawasha, near Salisbury [now Harare]; [17°46ʹ14ʺ S, 31°15ʹ11ʺ E]; Mar. 1978; A. Watsham leg.; BMNH GoogleMaps 1 ♀; same collection data as for preceding but Dec. 1978; BMNH GoogleMaps 1 ♀; same collection data as for preceding but Feb. 1979; BMNH GoogleMaps 1 ♂; same collection data as for preceding but Dec. 1981; BMNH. – GoogleMaps Mashonaland West • 2 ♂♂, 1 ♀; Lion and Cheetah Park 24 km W of Harare; 17°50ʹ S, 30°49ʹ E; 17–18 Dec. 1995; W.J. Pulawski leg.; CAS. – GoogleMaps Midlands • 1 ♀; Selukwe [now Shurugwi]; [19°40ʹ S, 30°00ʹ E]; 1911; A. Ellenberger leg.; MNHN GoogleMaps .

Description

Female

SIZE. 22.1–25.8 mm.

COLOR. Black except for basal half of mandible, which is ferruginous. Wings fuscous, with cyan-violet iridescence.

VESTITURE. Appressed setae on clypeus black, on paraocular area silvery with brown luster, on collar, scutum and propodeal enclosure black. Erect setae on clypeus, paraocular area, collar, scutum and propodeal enclosure black. Erect propodeal setae oriented perpendicularly, slightly curved towards anterior. Central and lower part of clypeus glabrous. Scutellum densely and coarsely pubescent.

STRUCTURE. Free clypeal margin medially with broad, notched tooth, stepped above. Clypeus without indentation or carina. Scutellum convex. Metanotum slightly raised, not notably bituberculate. 2 nd recurrent vein joins at interstitium between submarginal cells II and III. Propodeal enclosure without

any notable ridges. Foretarsomere I 2.7–3.6 × length of antepenultimate spine. Petiole length 1.9–2.5 × its medial width.

Male

SIZE. 19.6–24.3 mm.

COLOR. Black, except for ferruginous stripe in center of mandible. Wings fuscous, with cyan-violet iridescence.

VESTITURE. Appressed setae on clypeus and paraocular area silvery, on collar, scutum and propodeal enclosure black. Erect setae on clypeus, paraocular area, collar, scutum and propodeal enclosure black. Erect propodeal setae oriented perpendicularly, slightly curved towards anterior. Lower center of clypeus glabrous. Scutellum densely and coarsely pubescent.

STRUCTURE. Free clypeal margin medially with tooth emerging from inner side. Scutellum convex. Metanotum slightly raised, not notably bituberculate. 2 nd recurrent vein joins interstitium between submarginal cells II and III or slightly proximally. Propodeal enclosure without any notable ridges. Posterior margin of metasomal tergum VII convex. Posterior margin of metasomal sternum VII simple, of metasomal sternum VIII triangular. Petiole length 2.0–2.8× its medial width. Flagellomere V and often VI with moderately broad placoid covering their entire length and tapering toward base and apex.

Variation

Specimens from Lesotho and Gauteng Province ( var. basuto / kobrowi ) can vary as follows:

Female

COLOR. Black except for the following, which are ferruginous: basal half of mandible, lower part of clypeus, scape, pedicel, fore and midlegs from apical half of femur, hindleg from femur onward, excluding apical half of claw, and apical metasomal segment. Wings fuscous, with violet luster.

VESTITURE. Appressed setae on clypeus brassy-ferruginous, on paraocular area silvery, on collar, scutum and propodeal enclosure black. Erect setae on clypeus ferruginous near free margin, black otherwise, on paraocular area, collar, scutum and propodeal enclosure black. Erect propodeal setae oriented perpendicularly, slightly curved towards anterior. Lower center of clypeus glabrous. Scutellum densely and coarsely pubescent.

STRUCTURE. Free clypeal margin medially with broad, notched tooth, stepped above. Clypeus with indistinct indentation in lower center, without carina. Scutellum convex. Metanotum raised, not notably bituberculate. 2 nd recurrent vein joins slightly proximal from interstitium between submarginal cells II and III. Propodeal enclosure without any notable ridges. Foretarsomere I 2.8–2.9 × length of antepenultimate spine. Petiole length 1.70–1.83 × its medial width.

Male

COLOR. Black except for the following, which are ferruginous: basal half of mandible, femora apically, tibiae, and mid- and hindtarsomeres I–III. Wings hyaline, with yellowish tinge, apical margin of forewing infuscate.

VESTITURE. Appressed setae on clypeus and paraocular area silvery, on collar and scutum black, on propodeal enclosure brownish-gray. Erect setae on clypeus, paraocular area, collar, scutum and propodeal enclosure black. Erect propodeal setae oriented perpendicularly, slightly curved towards anterior. Lower center of clypeus glabrous. Scutellum densely and coarsely pubescent.

STRUCTURE. Free clypeal margin medially with tooth emerging from inner side. Scutellum convex. Metanotum slightly raised, not notably bituberculate. 2 nd recurrent vein nearly joins interstitium between submarginal cells II and III. Propodeal enclosure without any notable ridges. Posterior margin of metasomal tergum VII convex. Posterior margin of metasomal sternum VII simple, of metasomal sternum VIII triangular. Petiole length 1.5–1.6 × its medial width. Flagellomere V with moderately broad placoid covering its proximal four-fifths.

Distribution

Eastern, southeastern and southwestern Africa.

Remarks

As van der Vecht (1961) correctly stated, the name Sphex haemorrhoidalis Fabricius, 1781 , which Arnold (1928) used for black-legged specimens, is in fact linked to a red-legged taxon from western Africa. Other authors like Leclercq (1955, 1961), Berland (1956) and Dollfuss (2008) have based their identifications on Arnold’s mistake, and therefore many locality records and previous synonymizations were erroneous.

Although we were unable to examine the types of S. kobrowi (despite our request, it was apparently not included in the loan from TMP) or S. basuto , we nonetheless feel that our opinion on their synonymy is justified. First and foremost, we did study specimens which matched the descriptions and were most likely from the same collecting series as the types of both taxa, as they shared their locality, time frame and presumed collector. Labels indicated that Arnold had examined some of them as well. Secondly, we would have proposed to synonymize S. basuto with S. kobrowi even without additional evidence beyond their original descriptions. Judging from Arnold’s diagnosis, both resemble each other very much and are only distinguished through S. basuto having “thinner” propodeal pubescence and a slightly different interocular distance as well as flagellomere length. The measurements for the varieties of S. haemorrhoidalis in Arnold (1947) all seem quite dubious, especially in the case of S. basuto and S. kobrowi , with differences in length of less than 4% being used to differentiate between them. In addition, the actual numbers given are virtually useless for identifications without material for comparison, as they are highly dependent on the angle from which the measurements are taken and the how the boundaries between areas are perceived, yet these details are nowhere mentioned.

Though distinct in coloration and with a slightly different ratio of petiole length to width, S. kobrowi appears to be genetically identical to S. umtalicus . A comparison of parts of the CO1 sequences of specimens conforming to both taxa showed their complete congruence.

DEI

Germany, Muencheberg, Deutsches Entomologisches Institut im ZALF

MNHN

France, Paris, Museum National d'Histoire Naturelle

IRSN

Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique

BMNH

United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]

ZMB

Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet

MRAC

Belgium, Tervuren, Musee Royal de l'Afrique Centrale

AMNH

USA, New York, New York, American Museum of Natural History

ICIPE

ICIPE

NMK

Kenya, Nairobi, National Museum of Kenya

CAS

USA, California, San Francisco, California Academy of Sciences

NRS

Sweden, Stockholm, Naturhistoriska riksmuseet

TMP

Taiwan [Republic of China], Taipei, Taiwan Provincial Museum

RMNH

Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]

AMG

South Africa, Cape Province, Grahamstown, Albany Museum

OUMNH

United Kingdom, Oxford, University Museum of Natural History

DEI

Senckenberg Deutsches Entomologisches Institut

IRSN

Institut Royal des Sciences Naturelles de Belgique

ZMB

Museum für Naturkunde Berlin (Zoological Collections)

MRAC

Musée Royal de l’Afrique Centrale

AMNH

American Museum of Natural History

CAS

California Academy of Sciences

NRS

Naturhistoriska Riksmuseet

TMP

Transvaal Museum

RMNH

National Museum of Natural History, Naturalis

AMG

Albany Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hymenoptera

Family

Vespidae

Genus

Sphex

Loc

Sphex umtalicus Strand, 1916

Dörfel, Thorleif H. & Ohl, Michael 2022
2022
Loc

Chlorion haemorrhoidalis var. basuto

Arnold G. 1947: 146
1947
Loc

Chlorion haemorrhoidalis var. kobrowi

Arnold G. 1928: 367
1928
Loc

Sphex umtalicus

Strand E. 1916: 105
1916
Loc

Sphex nigripes var. pachyderma

Strand E. 1916: 106
1916