Clivina dostaliana, Balkenohl, 2018

Balkenohl, Michael, 2018, The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini), Linzer biologische Beiträge 50 (1), pp. 189-195 : 191-194

Clivina dostaliana nov.sp.

( Figs 2-4, 7-9 )

Type material: Holotype: ♂, with labels and data: blue, printed " NAMIBIA-Exp. ZMB 1992 Kavango: Gelukkie Kavango-Ufer , 18°03’S / 21°08’E. 1.III.92 leg. M. Uhlig " ( MFNB). Paratypes: 5♀♀: same data as holotype ( MFNB, CBB).

Etymology: The species is dedicated to Dr. Alexander Dostal (Vienna) and is expressed as Latinised adjective.

Diagnosis: A medium sized, cylindriform species, with a row of six short but big and a second row with smaller setae arising from tubercle-like punctures at the mesotibia, and small clypeal wings separated from middle part by small obtuse notches. Distinguished from the other species of the subgenus by the flattened eyes (dorsal view) and the reduced shape of the eye in lateral view, and by the supraantennal plates which are as broad as the eyes. Moreover, the shape of the median lobe of the aedeagus and the structure of the internal sac is different.

Description:

Measurements: Body length 2.85–2.95 mm (ẍ = 2.92 mm *), width 0.7–0.72 mm (ẍ = 0.71 mm *), ratio length/width of pronotum 1.19–1.24 (ẍ = 1.21*), ratio length/width of elytra 2.26–2.42 (ẍ = 2.32*); (*n = 6).

Colour: Shiny. Red-brown. Legs, antennae and mouthparts yellow, front femur brown, apex and margin of mandible darker, supraantennal plates translucent yellowish.

Head ( Figs 3, 4 ): Less than a quarter narrower than pronotum. Clypeus straight anteriorly, wings small, not as advanced as clypeus, separated from clypeus by small obtuse notches; supraantennal plates wide, margins convex; clypeus, wings, and supraantennal plates with small reflexed margin. Supraantennal plates with carina posteriorly extending up to mideye level, anteriorly separated from wings by obtuse-angled notches. Frons with distinct carina in form of an inverted "V", prolonged anteriorly as keel, with second but curved carina anteriorly, with some longitudinal wide rugae between carinae and on posterior part of frons. Supraorbital carinae relatively short, separated from eyes by supraorbital furrows, each with two supraorbital setae. Neck laterally with few punctures of moderate size. Eyes distinctly flattened, as high as supraantennal plates, shape in lateral view is remindful of a truncated pyramid. Genae indistinct. Antenna short, just reaching mid-length of pronotum, segments five to ten wider than long. Labrum straight anteriorly, with irregular reticulation, five-setose. Mandible moderately curved, also towards apex, arcuate at apex, finely margined in apical half. Apical segment of maxillary palpomere securiform, pointed at apex, terminal segment of labial palpomere bottle like, slender apically. Mentum with median tooth developed a pointed slender knob, not as projecting anteriorly as lateral lobes. Lateral lobes finely carinate, surface with irregular reticulation.

Pronotum: Slightly convex on disc (lateral view). Cylindrical, longer than wide. Lateral margin nearly straight in dorsal view, sigmoid in lateral view. Anterior angles rounded off by reflexed lateral margin. Reflexed lateral margin smooth. Lateral channel narrow, slightly widened at level of anterior setigerous punctures. Anterior and posterior setigerous punctures situated in marginal channel. Posterior angle not visible. Pleura visible in dorsal view. Anterior margin excised. Anterior transverse line distinct, not reaching anterior angles. Median line sharp, running over level of anterior transverse line but not joining, nearly reaching anterior margin. Distinctly joining basal constriction. Basal constriction moderately wide, flange with sharp carina. Disc usually without bigger punctures (in two paratypes unilaterally with few paralateral punctures), with indistinct longitudinal basal impression bilaterally. Surface with transverse wrinkles.

Elytra: Cylindrical, two and a third times longer than wide. Marginal channel running over rounded humerus up to level of third stria without joining striae at base, with umbilical setigerous punctures widely interrupted at middle, thought two to three punctures are missing. Reflexed margin smooth. No humeral tooth. Striole missing, basal tubercle with setigerous puncture distinct, situated at declivity of first stria, Striae one and two free at base, all striae deep, punctuate-striate, stria one joining stria seven at apex, stria two ending free at apex, striae three and four and five and six joining at apex. Intervals distinctly convex, seventh carinate apically. No setigerous punctures on interval three. Surface shiny, with some indistinct wrinkles.

Hind wings: Fully developed.

Lower surface: Pleura distinctly swollen postero–laterally, with transverse wrinkles and isodiametric reticulation (160 times). Proepisternum and abdominal sternite covered with isodiametric to imbricate reticulation. Basal abdominal sternites with isodiametric reticulation, last three sternites shiny, with few microscopic punctures. Abdominal sternum with the two marginal setae widely separated. No differences observed among sexes.

Legs: Protibia with strong digitation, with two big, one smaller and one minute lateral preapical denticles, surface with longitudinal reticulation, apical spine as long as spur, movable spur ensiform, acute at apex, tarsomeres sturdy. Mesotibia without setae on the inner side, dorso-laterally with row of six big and short setae arising from tubercle-like punctures, preapical seta furnished spine missing.

Male genitalia ( Fig 7, 8 ): Median lobe moderately slender, moderately arcuate in dorsal view, slightly more curved at middle, nearly straight in lateral view, tip of apex rounded like an elongated spoon. Surface of apical third dorsally with over a dozen of fine striae pointing apically. Internal sac at the beginning of apical third with 6 teeth pointing basally, basal sclerite distinct, V-shaped. Parameres conspicuously asymmetrical, somewhat distorted, each with one short preapical seta.

Female genitalia ( Fig 9 ): Coxostyli thin, hyaline. Terminal one like a flattened club, with two setae in the middle part. Basal one broad, velum-like.

Variation: Variation noted on the pronotum which exhibits in two of the paratypes unilaterally few paralateral punctures. In other specimens the basal impression is more or less well visible, and the length of the median line on the pronotum differs after passing through the anterior transverse line.

Distribution: Known from the banks of the Kavango River in the North of Namibia.

Key to the species

1 Eyes convex in dorsal view, supraantennal plates not as wide as eyes ( Fig. 1 ). Pronotum bilaterally with distinct paramedian and/or paralateral row or cloud-like group of punctures. Frons of head more or less smooth between the carinae ............................... 2

- Eyes flattened in dorsal view ( Fig. 2, 3 ), supraantennal plates as wide as eyes, shape in lateral view like a truncated pyramid ( Fig. 4 ). Pronotum without paramedian and/or paralateral punctures. Frons of head with longitudinal wide rugae between the carinae. Only 2-3 punctures laterally at neck level. V-like carina on the frons of head reaching posteriorly nearly up to neck level. Length 2.9 ................................... dostaliana nov.sp.

2 Eyes in lateral view semicircular ( Fig. 6 ). Lateral channel of pronotum slightly widened at level of anterior setigerous puncture. Carinae on the frons of head robust, V-like carina prolonged over posterior eye level. ................................................................................ 3

- Eyes in lateral view circular ( Fig. 5 ). Lateral channel of pronotum distinctly widened at level of anterior setigerous puncture. Carinae on frons of head fine, V-like carina reaching up to posterior eye level. Length 2.5-2.7 mm ................ physopleura BURGEON

3 Pronotum with group of longitudinal paramedian and paralateral punctures. Angle of Vlike carina on frons anteriorly not sharp, somewhat rounded. With distinct group of punctures at neck level but interrupted at middle. Length 3.0 mm....... bulirschi DOSTAL

- Pronotum with group of longitudinal paralateral punctures. Angle of V-like carina on frons anteriorly truncated. Group of punctures at neck level nearly uninterrupted at middle. Length 3.3 mm ............................................................. donabaueriana DOSTAL

Figures

Figs 1-9: Clivina (subgenus Physoclivina) spp.: (1) C. physopleura BURGEON, holotype, habitus; (2) C. dostaliana nov.sp., holotype, habitus; (3-6) head and pronotum: (3) C. dostaliana nov.sp., dorsal view; (4) lateral view; (5) C. physopleura BURGEON, lateral view; (6) C. bulirschi DOSTAL, lateral view; (7) C. dostaliana nov.sp., holotype, aedeagus and parameres, dorsal view and (8) lateral view; (9) C. dostaliana nov.sp., female coxostyli.