Heteroschizomus goodnightorum Rowland, 1973

Monjaraz-Ruedas, Rodrigo, Prendini, Lorenzo & Francke, Oscar F., 2019, Systematics Of The Short-Tailed Whipscorpion Genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), With Descriptions Of Six New Genera And Five New Species, Bulletin of the American Museum of Natural History 2019 (435), pp. 1-93 : 47-48

Heteroschizomus goodnightorum Rowland, 1973

Figures 4 , 6, 8E, 21A–C; table 7

Heteroschizomus goodnightorum Rowland, 1973a: 2–6 ; 1973b: 202; 1973c: 136; 1975b: 31; Rowland and Reddell, 1977: 83; Reddell and Cokendolpher, 1991: 18; 1995: 3, 5, 101, 106.

Schizomus goodnightorum: Rowland 1975a: 34 , 167, 168, 194, 257–269, 325, 397; Rowland and Reddell, 1977: 90, 99, 100; 1979a: 163, 164; 1981: 19, 21–27; Reddell and Cokendolpher, 1995: 3, 106.

Stenochrus goodnightorum: Reddell and Cokendolpher, 1991: 18 ; 1995: 5, 8, 18, 102, 106; Villarreal et al., 2008: 64; Villarreal and García, 2012: 5; Monjaraz-Ruedas and Francke, 2016: 783, 784, 797, 804; 2018: 190.

TYPE MATERIAL: MEXICO: Yucatan: Chichén Itzá, vi.1948, C. Goodnight, holotype ♂, paratype ♂ ( AMNH).

SUPPLEMENTARY DESCRIPTION: The following description of the female is based on the additional material examined.

Prosoma: Propeltidium with two apical setae; three pairs of dorsal setae.

Chelicerae: Movable finger with very small lamella; serrula with 13 teeth, guard tooth present. Fixed finger with five small teeth between two primary teeth; setal group formula, 3-6-4-2-7-8-1-6.

Pedipalps: Pedipalps homeomorphic; 1.59× longer than propeltidium. Trochanter with small rounded apical process, not projected; prolateral surface with small apical spur. Femur 1.73× longer than high; retroventral margin with setae Fe 1, Fe 5, Fev 1, and Fev 2 acuminate; prolateral surface with row of three ventral setae (Fmv 1–3) and two dorsal setae (Fmd 2, Fmd 3). Patella with three acuminate Pe setae and four feathered Pm setae; without distinctive armature. Tibia setal formula, 3:3:4; Ter acuminate, Tmr and Tir feathered. Tarsal spurs asymmetric.

Opisthosoma: Segments X–XII not elongated.

Pygidial flagellum: Flagellum (♀) with three flagellomeres (fig. 21A– C); seta Dl 2 reduced, situated posterior to Vl 1, Dl 3 situated posterior to Vl 2, Vm 2 present, Vm 1 aligned with Vm 2; Dl 1 and Dl 4 microsetae present.

Female spermathecae: Two pairs of lobes (fig. 8E); median and lateral lobes conical, subequal in length and width; median lobes linear, lateral lobes sublinear, apex slightly curved laterally, unsclerotized apically, with few duct openings; median lobe bases aligned with lateral lobe bases. Chitinized arch mug shaped; anterior branch not sclerotized. Gonopod wide; width ca. 2×length.

REMARKS: Rowland and Reddell (1977) synonymized Heteroschizomus , stat. rev., and transferred H. goodnightorum to Schizomus , based on the discovery of other species with an elongated opisthosoma. Reddell and Cokendolpher (1991) transferred Schizomus goodnightorum to Stenochrus based on the morphology of the female spermathecae described by Rowland and Reddell (1981). The discovery and correct identification of the female of H. goodnightorum confirmed the validity of Heteroschizomus and the correct placement of H. goodnightorum and related species, discoveries that were independently supported by the DNA sequence data.

ADDITIONAL MATERIAL EXAMINED: MEX- ICO: Yucatán: Municipio de Chichén Itzá: Chichen Itzá , vi.1948, C. Goodnight, 1 ♀ ( AMNH) ; Fuera de la Cueva de Kanthon, 20°38′08″N 88°29′42″W, 23 m, 2.viii.2014, J. Mendoza and R. Monjaraz, 4 ♂ (AMCC [LP 14522]), 2 ♂, 1 ♀ (CNAN Sz171). Municipio de Kaua: Fuera de la Cueva de Kaua , 20°37′04″N 88°25′26″W, 24 m, 5.x.2014, G. Contreras and G. Montiel, 1 ♂, 2 ♀ (CNAN Sz163). Quintana Roo: Municipio de Cancun : Km 234 Highway Mérida – Cancun, 20°53′57″N 87°28′25″W, 19 m, 20. vii.2010, D. Barrales, G. Contreras, J. Cruz, O. Francke, G. Montiel, M. Paradiz, C. Santibañez, and A. Valdez, 1 ♂ (CNAN Sz8) .

Figures

FIG. 4. Map of southern Mexico, plotting known locality records for the short-tailed whipscorpion genera Heteroschizomus Rowland, 1973, stat. rev. (circles), and Troglostenochrus, gen. nov. (squares) (Schizomida: Hubbardiidae Cook, 1899), based on data from museum collections and the literature.