Malagiella valterova, Ubick & Griswold, 2011

Ubick, Darrell & Griswold, Charles E., 2011, The Malagasy Goblin Spiders Of The New Genus Malagiella (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2011 (356), pp. 1-86 : 78-79

Malagiella valterova , new species

Figures 236–267 , 473 , 483 , 487 ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifted litter along Tatamaly River, 21 ° 309420S, 47 ° 249360E, 1075 m, 2 km W Andrambovato, Fianarantsoa Province, Madagascar (3–5 Jun 2005, B. Fisher), deposited in CAS (CASENT 9030812, PBI _OON 03461).

ETYMOLOGY: This species is named after Darrell Ubick’s late father, ‘‘Valter’’ Ubick, in remembrance of many years of companionship, and is given in Croatian genitive.

DIAGNOSIS: The male of this species most closely resembles M. ranomafana , from which it differs by its longer palpal patella and bulbcymbium (fig. 483), and by the slightly longer ventral abdominal scute (fig. 239). The female differs from others in the group in having a widely sinuous receptaculum, W/L ca. 0.7 (figs. 264–267).

MALE (PBI_OON 03461): Total length 1.55, carapace length 0.78, width 0.62. CEPHALOTHORAX: Carapace as in M. ranomafana (figs. 237, 238, 241, 242). Clypeus as in M. ranomafana , but slightly longer (figs. 238, 241). Eyes as in M. ranomafana , eye area width 0.5 carapace width; length subequal to clypeus length. Sternum as in M. ranomafana (figs. 240, 473). ABDOMEN (figs. 239, 243–245) as in M. ranomafana . ABDOMEN: Ventral scute (fig. 239) slightly longer than in M. ranomafana . GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 263). Palp (figs. 256–262, 483) slightly larger than in M. ranomafana .

FEMALE (PBI_OON 03461): Total length 1.70, carapace length 0.80, width 0.66. CEPH- ALOTHORAX (figs. 246–248, 251, 252): Clypeus shorter than eye region length (fig. 248). Eye area 1.6 clypeus length (figs. 247, 248, 251). ABDOMEN (figs. 249, 253– 255): Dorsal scutum slightly longer than in M. ranomafana . GENITALIA: Ventral view: receptaculum slender and strongly sinuous, W/L about 0.7 (figs. 264–267, map 4).

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality.

Figures

Figs. 236–245. Malagiella valterova, new species, male (PBI_OON 03461). 236. Habitus, ventral view. 237. Same, dorsal view. 238. Same, lateral view. Scale bar 5 1 mm. 239. Abdomen, ventral view. 240. Cephalothorax, ventral view. 241. Same, anterior view. 242. Same, posterior view. 243. Abdomen, anterior view. 244. Same, lateral view. 245. Same, posterior view.

Figs. 246–255. Malagiella valterova, new species, female (PBI_OON 03461). 246. Habitus, ventral view. 247. Same, dorsal view. 248. Same, lateral view. Scale bar 5 1 mm. 249. Abdomen, ventral view. 250. Cephalothorax, ventral view. 251. Same, anterior view. 252. Same, posterior view. 253. Abdomen, anterior view. 254. Same, lateral view. 255. Same, posterior view.

Figs. 256–263. Malagiella valterova, new species, male (PBI_OON 03461). 256. Palp, femur to bulb, prolateral view. 257. Same, retrolateral view. 258. Same, dorsal view. 259. Same, ventral view. 260. Same, anterior view. 261. Same, posterior view. 262. Cephalothorax, sublateral view, showing embolus in resting position (arrow). 263. Abdomen, sublateral view, showing sperm pore (arrow).

Figs. 264–267. Malagiella valterova, new species, female (PBI_OON 03461). 264. Abdomen, ventral view. 265. Same, more posterior view. 266. Epigynal region, ventral view. 267. Same, lateral view.

Figs. 472–480. Malagiella species, sternum, ventrolateral view, males (472–477) and females (478–480). 472. M. andringitra, PBI_OON 03234. 473. M. valterova, PBI_OON 03461. 474. M. fisheri, PBI_OON 03452. 475. M. ranomafana, PBI_OON 01999. 476. M. vohiparara, PBI_OON 03078. 477. M. toliara, PBI_OON 03377. 478. M. ranomafana, PBI_OON 01999. 479. M. vohiparara, PBI_OON 03078. 480. M. toliara, PBI_OON 03221.

Figs. 481–486. Malagiella species, male palpus, femur to bulb, prolateral view. 481. M. ranomafana, PBI_OON 03224. 482. M. andringitra, PBI_OON 03234. 483. M. valterova, PBI_OON 03461. 484. M. vohiparara, PBI_OON 03078. 485. M. fisheri, PBI_OON 03452. 486. M. toliara, PBI_OON 03377. Scale bar (200 Mm) applies to all palpi.

Fig. 487. Size variation in Malagiella measured as a function of carapace length to carapace width. Plot is based on measurements of 26 male specimens (‘‘x’’) and 30 female specimens (‘‘o’’). Blue lines enclose male values for a given species and red lines female values; disjunct conspecific readings are connected by straight lines. Green dotted lines enclose the three species groups. Brown dashed lines refer to the size of female DS, with the large dashes (upper right) enclosing species with the largest scutes and small dashes (lower left) those with the smallest. Abbreviations for species names: amba 5 M.ambalavo, andr 5 M. andringitra, fish 5 M. fisheri, good 5 M. goodmani, niki 5 M. nikina, rana 5 M. ranavalona, rano 5 M. ranomafana, toli 5 M. toliara, valt 5 M. valterova, vohi 5 M. vohiparara.