Malagiella ranomafana, Ubick & Griswold, 2011Malagiella valterova, Ubick & Griswold, 2011Malagiella andringitra, Ubick & Griswold, 2011Malagiella fisheri, Ubick & Griswold, 2011

Ubick, Darrell & Griswold, Charles E., 2011, The Malagasy Goblin Spiders Of The New Genus Malagiella (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2011 (356), pp. 1-86: 59-80

publication ID

http://doi.org/ 10.1206/356.1

persistent identifier

http://treatment.plazi.org/id/03C2C67F-FFD4-FF84-F0FD-F99FFBC7FC29

treatment provided by

Tatiana

scientific name

Malagiella ranomafana Malagiella valterova Malagiella andringitra Malagiella fisheri
status

new species

Malagiella ranomafana   , new species

Figures 7–116 View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs View Figs , 200–235 View Figs View Figs View Figs View Figs , 463, 466, 467 View Figs , 475, 478 View Figs , 481 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype, female allotype, and female paratype from Talatakely, P.N. Ra- nomafana, 21 ° 159S, 47 ° 259E, 900 m, Fianarantsoa Province, Madagascar (5–7 Dec 1993, N. Scharff, S. Larcher, C. Griswold, R. Andriamasimanana), deposited in CAS (CASENT 9029645, PBI _ OON 03225 View Materials ).

ETYMOLOGY: The species is named after the type locality.

DIAGNOSIS: The male may be distinguished from other species in the group by the combination of sternum strongly bristly, bulb-cymbium small (about 0.5 patella-tibia length), and a palpal patella of moderate size (fig. 481). The female differs by the combination of large dorsal scutum, PES with straight posterior margin, and receptaculum moderately sinuous (W/L 5 0.3–0.5) (figs. 228–235).

MALE (PBI_ OON 03225 View Materials ): Total length 1.70 (1.56–1.70), carapace length 0.80 (0.78– 0.80), width 0.64 (0.62–0.66), N 5 12. CEPHALOTHORAX: Carapace ovoid in dorsal view, pars cephalica slightly elevated in lateral view, anteriorly narrowed to about 0.55 times its maximum width; lateral margin strongly undulate (fig. 14). Eye area width 0.5 carapace width, length 1.2 clypeus length (figs. 13–15). Sternum lateral margin infracoxal grooves present, with anterior and posterior openings (fig. 19). Mouthparts: Cheliceral paturon posterior surface with a row of three long setae (fig. 19); with promarginal tooth and denticles on both margins (fig. 25). Endite with median lobe bearing dense scopula and lateral lobe with serrula and three stout setae (figs. 26, 27). ABDOMEN: Pedicel tube of medium length (fig. 208), dorsal extension of epigastric scute about one pedicel diameter (fig. 207). Spinnerets: ALS with three subequal spigots; PMS with two subequal spigots; PLS with four subequal spigots (figs. 40–43). LEGS: Tarsi I–IV superior claws examined in detail: tarsal proclaws and retroclaws with faces striate; tarsi I, II superior claws with three large teeth on lateral surfaces and three small teeth on median surfaces of both proclaw and retroclaw; tarsi III, IV superior claws with three teeth on lateral surfaces of proclaw and retroclaw, lacking teeth on median surfaces; tarsi I–IV lacking inferior claw (figs. 74–78); tarsal claws III, IV each with three modified clawlike setae (figs. 77, 78). Trichobothria examined with SEM: trichobothrial base longitudinally narrowed, aperture internal texture not gratelike; hood covered by numerous low closely spaced ridges, usually in transverse orientation but radially arranged in proximal half of some bothria; both types of ridges occur on tibial trichobothria (figs. 62–64). Tarsal organ with 2 sensilla on palp and tarsi III, IV; 3 sensilla on tarsi I, II (figs. 57–61). GENITALIA: Epigastric region with sperm pore small, rebordered (figs. 34–36). Palp (figs. 85–97, 220–227, 481, map 3) bulb pale orange, more than 2 times as long as cymbium; embolar complex (figs. 93–97) distal, compressed, retrolaterally concave, with hyaline ventral prong (BVP) and dorsal embolus, apex transversely expanded with median opening and lateral prongs, prolateral prong (EPP) with groove connecting embolar opening (fig. 98).

FEMALE (PBI_ OON 01999 View Materials ): Total length 1.71 (1.71–1.96), carapace length 0.82 (0.80– 0.84), width 0.66 (0.64–0.68), N 5 11. CEPHALOTHORAX: Clypeus shorter than eye region length (fig. 18), anteriorly narrowed to about 0.49 times its maximum width. Eyes (figs. 16–18, 215) as in male. Sternum microsculpture covering entire surface (figs. 21–23). Chelicerae (figs. 28, 29) and labium (fig. 29) as in male. Endites lacking anteromedian projection and anterolateral knob, three setae near serrula not stout (figs. 29, 30). ABDOMEN: Pedicel tube of medium length (fig. 218) and epigastric scutum dorsal extension (fig. 217), both slightly shorter than in male. Dorsal scutum covering more than 0.7 of abdomen (figs. 211, 212), not fused to epigastric scutum (figs. 37, 38, 217, 218). Postepigastric scutum short, extending only to posterior spiracles, with small lateral sclerites, internally with short posteriorly directed lateral apodemes (figs. 108, 116, 213). Spinnerets: ALS with three spigots, median slightly larger; PMS with three subequal spigots; PLS with six subequal spigots (figs. 44–47). LEGS (figs. 466, 467): With fine transverse ridges delineating regions of muscle attachment (figs. 54, 55) and slit sensilla (figs. 51–53). Tarsal claws as in male except: tarsus IV superior claws with three teeth on lateral surfaces and one large apical tooth on median surfaces of both proclaw and retroclaw (figs. 79–84). Trichobothria (figs. 70–73) as in male. Tarsal organ (figs. 65–69) as in male. GENITALIA: Ventral view (figs. 100–107, 228–234, map 4): epigynal area with median depression enclosing small round hole (copulatory opening) (figs. 101, 103, 105), with a pair of slitlike openings at base of apodemes (fig. 104), with sigmoid tube (receptaculum) and a pair of paramedian apodemes visible through cuticle (figs. 228, 234). Dorsal view (figs. 108–116, 235): receptaculum sigmoid, reaching posteriorly slightly beyond postepigastric scutum; with pair of lateral apodemes. The receptaculum varies considerably in size, degree of sinuation, and direction of curvature (figs. 228–235).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: Ranomafana N.P., 21 ° 129S, 47 ° 279E, forest, 1–30 Apr 1992 (V. Roth, B. Roth, S. Kariko, CASENT 9029651, PBI _ OON 03231 View Materials ), 1♀   ; Ranomafana N.P., Maharira , 09 Apr 1992 (Emile, Kariko-Roth Coll., MCZ 53653, MCZ 53656, PBI _ OON 00028216, PBI _ OON 00028217), 2♀   ; Trail F, 325 m, 18 May 1992 (Emile, Kariko-Roth Coll., MCZ 53652 View Materials , PBI _ OON 00028212), 1♀   ; Trail L, 1050 m, 15 Apr 1992 (Emile, Kariko-Roth Coll., MCZ 53626 View Materials , PBI _ OON 00028214), 2♀   ; Trail M, 15 May 1992 (Albert, Kariko- Roth Coll., MCZ 53657 View Materials , PBI _ OON 00028215), 1♀   ; across footbridge at campsite, 26 Mar 1992 (S. Kariko, V. Roth, MCZ 53646 View Materials , PBI _ OON 00028213) 6♀   ; Talatakely, P.N. Ranomafana , 21 ° 14.99S, 47 ° 25.69E, sifting leaf litter, 5–18 Apr 1998 (C. Griswold, D. Ubick, CASENT 9029647, PBI _ OON 03227 View Materials ), 18, 2♀ GoogleMaps   ; 19–30 Apr 1998 (C. Griswold, D. Ubick, CASENT 9029649, PBI _ OON 03229 View Materials ), 1♀   ; pitfall trap, 13–27 Apr 1998 (Griswold et al., CASENT 9029650, PBI _ OON 03230 View Materials ), 18   ; 21 ° 15.39S, 47 ° 25.99E, pitfall trap, 9–26 Apr 1998 (Griswold et al., CASENT 9029644, PBI _ OON 03224 View Materials ), 28, 3♀ GoogleMaps   ; 21 ° 159S, 47 ° 269E, 915– 1000 m, 30 Oct–20 Nov 1998 (V. Lee, K. Ribardo, CASENT 9029648, PBI _ OON 03228 View Materials ), 18   ; 21 ° 15.89S, 47 ° 25.29E, 940– 985 m, pitfall trap, 1–21 Nov 1998 (V. Lee, K. Ribardo, CASENT 9029646, PBI _ OON 03226 View Materials ), 18 GoogleMaps   ; 21 ° 15.59S, 47 ° 25.4 E, 1050 m, montane rainforest, 24 Dec 2005 – 14 Jan 2006 (H. Wood, J. Miller, et al., HW009, CASENT 9019096, PBI _ OON 03433 View Materials ), 28, 2♀ GoogleMaps   ; Vatoharanana 1100, 221.29, 47.43333, 1100 m, montane rainforest, pitfall trap, 27– 31 Mar 2003 (Fisher, Griswold et al., BLF8402, CASENT 9011546, PBI _ OON 03232 View Materials ), 28, 1♀   ; sifted litter, leaf mold, rotten wood, 27–31 Mar 2003 (Fisher, Griswold et al., BLF8400, CASENT 9011421, PBI _ OON 02216 View Materials ), 1♀   ; Vatoharanana 1200, 21 ° 16.79S, 47 ° 26.19E, 1200 m, primary rainforest understory, 15–29 Apr 1998 (Griswold et al., CASENT 9025683, PBI _ OON 01999 View Materials ), 48, 3♀ GoogleMaps   ; sifting leaf litter, 15 Apr 1998 (C. Griswold, D. Ubick, CASENT 9029652, PBI _ OON 03233 View Materials ), 1♀   ; 7 km SW Ranomafana , 21.126N, 47.276E, 1000 m, 20–24 Mar 1990 (W. Steiner, USNM, PBI _ OON 00027793), 1♀ GoogleMaps   .

DISTRIBUTION: Known only from Ranomafana National Park, Madagascar.

Malagiella valterova   , new species

Figures 236–267 View Figs View Figs View Figs View Figs , 473 View Figs , 483 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifted litter along Tatamaly River, 21 ° 309420S, 47 ° 249360E, 1075 m, 2 km W Andrambovato, Fianarantsoa Province, Madagascar (3–5 Jun 2005, B. Fisher), deposited in CAS (CASENT 9030812, PBI _ OON 03461 View Materials ).

ETYMOLOGY: This species is named after Darrell Ubick’s late father, ‘‘Valter’’ Ubick, in remembrance of many years of companionship, and is given in Croatian genitive.

DIAGNOSIS: The male of this species most closely resembles M. ranomafana   , from which it differs by its longer palpal patella and bulbcymbium (fig. 483), and by the slightly longer ventral abdominal scute (fig. 239). The female differs from others in the group in having a widely sinuous receptaculum, W/L ca. 0.7 (figs. 264–267).

MALE (PBI_ OON 03461 View Materials ): Total length 1.55, carapace length 0.78, width 0.62. CEPHALOTHORAX: Carapace as in M. ranomafana   (figs. 237, 238, 241, 242). Clypeus as in M. ranomafana   , but slightly longer (figs. 238, 241). Eyes as in M. ranomafana   , eye area width 0.5 carapace width; length subequal to clypeus length. Sternum as in M. ranomafana   (figs. 240, 473). ABDOMEN (figs. 239, 243–245) as in M. ranomafana   . ABDOMEN: Ventral scute (fig. 239) slightly longer than in M. ranomafana   . GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 263). Palp (figs. 256–262, 483) slightly larger than in M. ranomafana   .

FEMALE (PBI_ OON 03461 View Materials ): Total length 1.70, carapace length 0.80, width 0.66. CEPH- ALOTHORAX (figs. 246–248, 251, 252): Clypeus shorter than eye region length (fig. 248). Eye area 1.6 clypeus length (figs. 247, 248, 251). ABDOMEN (figs. 249, 253– 255): Dorsal scutum slightly longer than in M. ranomafana   . GENITALIA: Ventral view: receptaculum slender and strongly sinuous, W/L about 0.7 (figs. 264–267, map 4).

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality.

Malagiella andringitra   , new species

Figures 268–300 View Figs View Figs View Figs View Figs , 472 View Figs , 482 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype sifted from leaf litter in rainforest, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, Res. Andringitra, Fianarantsoa Province, Madagascar (23 Oct 1993, B. Fisher), deposited in CAS (8 holotype: CASENT 9029653, PBI _ OON 03234 View Materials ; ♀ allotype CASENT 9029669, PBI _ OON 03376 View Materials )   .

ETYMOLOGY: The species is named after the type locality.

DIAGNOSIS: The male differs from others in the group by the large bulb-cymbium (about 0.7 patella-tibia length; but about 0.5 in other species) and the presence of a shallow dorsal groove proximally on the bulb (fig. 482). The female differs in having a smaller dorsal scute (fig. 279) and the epigynal region weakly sclerotized and the receptaculum short and relatively straight (W/L about 0.25) (figs. 296, 297).

MALE (PBI_ OON 03234 View Materials ). Total length 1.53, carapace length 0.74, width 0.62. CEPHALOTHORAX: Carapace (figs. 269, 270, 274) as in M. ranomafana   . Eyes as in M. ranomafana   , but slightly smaller; eye area about 0.5 carapace width, length subequal to clypeus length (figs. 269, 273). Sternum (figs. 272, 472) as in M. ranomafana   . ABDO- MEN (figs. 269, 271, 275–277) as in M. valterova   , but with slightly smaller book lung covers. GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 271). Palp with bulb-cymbium large, about 0.7 patellatibia length; bulb with dorsal depression (figs. 288–295, 482, map 3).

FEMALE (PBI_ OON 03376 View Materials ). Total length 1.82 (1.80–1.82), carapace length 0.78, width 0.68 (0.66–0.68), N 5 2. CEPHALOTHO- RAX (figs. 278–280, 283, 284): Clypeus sloping forward in lateral view, shorter than eye row length (fig. 280). Eye area about 1.3 clypeus length (figs. 280, 283). ABDOMEN (figs. 281, 285–287): Pedicel tube short (fig. 286), scutum dorsal extension slightly less than pedicel diameter (fig. 286). Dorsal scutum smaller than carapace, covering about 0.6 abdomen length and width (fig. 279). Postepigastric scutum weakly sclerotized. LEGS: patella plus tibia I longer than carapace (fig. 280). GENITALIA: Epigynal region weakly sclerotized, receptaculum short and relatively straight, W/L about 0.25 (figs. 296–300, map 4).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: Res. Andringitra, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, rainforest, sifting leaf litter, 23 Oct 1993 (B. Fisher, CASENT 9029654, PBI_ OON 03375 View Materials ), 1♀.

DISTRIBUTION: Known only from the type locality in Res. Andringitra, Madagascar.

Malagiella fisheri   , new species Figures 301–328 View Figs View Figs View Figs , 474 View Figs , 485 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifting leaf litter in rainforest, at 28.5 km S Midongy-Sud, Mount Papango, 23 ° 50.19S, 46 ° 57.89E, 940 m, P.N. Befotaka-Midongy, Fianarantsoa Province, Madagascar (13–15 Nov 2006, Fisher et al.), deposited in CAS (CASENT 9030777, PBI _ OON 03452 View Materials ).

ETYMOLOGY: This species is named after Brian Fisher, collector of this and several other species of Malagiella   , in honor of his impressive sampling program of the Malagasy fauna.

DIAGNOSIS: The male of this species differs from other Malagiella   in lacking sternal bristles (fig. 474) and in having a huge palpal patella (figs. 321–324, 485). The female differs from others in having a distinctly procurved posterior margin of the PES (figs. 327, 328) and a slightly curved receptaculum (W/L about 0.3).

MALE (PBI_ OON 03452 View Materials ): Total length 1.62, carapace length 0.82, width 0.69. CEPHALOTHORAX: Eye area about 0.5 carapace width; subequal to clypeus length (figs. 302, 303, 306). Sternum lacking stiff bristles found in other species (figs. 305, 474), median concavity absent. ABDOMEN (figs. 304, 308–310): as in M. valterova   . GENITALIA (figs. 321–326): Palp with patella greatly enlarged (fig. 485), about five times femur length, but less than four times in other species (figs. 481–484, 485).

FEMALE (PBI_ OON 03452 View Materials ): Total length 1.70 (1.70–1.86), carapace length 0.80 (0.78– 0.80), width 0.70 (0.68–0.70), N 5 2. CEPHALOTHORAX (figs. 311–313): Clypeus shorter than eye region length. Eye area about 1.8 clypeus length (fig. 313). ABDO- MEN (figs. 314, 318–320): Pedicel tube (fig. 337) shorter than in male. GENITA- LIA: Postepigastric scutum with posterior margin procurved, receptaculum only slightly curved, W/L about 0.3 (figs. 327, 328).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: P.N. Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23 ° 50.59S, 46 ° 57.59E, 1250 m, montane rainforest, sifting leaf litter, 17–19 Nov 2006 (B. Fisher et al., CASENT 9030776, PBI_ OON 03442 View Materials ), 1♀.

DISTRIBUTION: Known only from P.N. Befotaka-Midongy, Madagascar.

Malagiella valterova   , new species

Figures 236–267 View Figs View Figs View Figs View Figs , 473 View Figs , 483 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifted litter along Tatamaly River, 21 ° 309420S, 47 ° 249360E, 1075 m, 2 km W Andrambovato, Fianarantsoa Province, Madagascar (3–5 Jun 2005, B. Fisher), deposited in CAS (CASENT 9030812, PBI _ OON 03461 View Materials ).

ETYMOLOGY: This species is named after Darrell Ubick’s late father, ‘‘Valter’’ Ubick, in remembrance of many years of companionship, and is given in Croatian genitive.

DIAGNOSIS: The male of this species most closely resembles M. ranomafana   , from which it differs by its longer palpal patella and bulbcymbium (fig. 483), and by the slightly longer ventral abdominal scute (fig. 239). The female differs from others in the group in having a widely sinuous receptaculum, W/L ca. 0.7 (figs. 264–267).

MALE (PBI_ OON 03461 View Materials ): Total length 1.55, carapace length 0.78, width 0.62. CEPHALOTHORAX: Carapace as in M. ranomafana   (figs. 237, 238, 241, 242). Clypeus as in M. ranomafana   , but slightly longer (figs. 238, 241). Eyes as in M. ranomafana   , eye area width 0.5 carapace width; length subequal to clypeus length. Sternum as in M. ranomafana   (figs. 240, 473). ABDOMEN (figs. 239, 243–245) as in M. ranomafana   . ABDOMEN: Ventral scute (fig. 239) slightly longer than in M. ranomafana   . GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 263). Palp (figs. 256–262, 483) slightly larger than in M. ranomafana   .

FEMALE (PBI_ OON 03461 View Materials ): Total length 1.70, carapace length 0.80, width 0.66. CEPH- ALOTHORAX (figs. 246–248, 251, 252): Clypeus shorter than eye region length (fig. 248). Eye area 1.6 clypeus length (figs. 247, 248, 251). ABDOMEN (figs. 249, 253– 255): Dorsal scutum slightly longer than in M. ranomafana   . GENITALIA: Ventral view: receptaculum slender and strongly sinuous, W/L about 0.7 (figs. 264–267, map 4).

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality.

Malagiella andringitra   , new species

Figures 268–300 View Figs View Figs View Figs View Figs , 472 View Figs , 482 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype sifted from leaf litter in rainforest, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, Res. Andringitra, Fianarantsoa Province, Madagascar (23 Oct 1993, B. Fisher), deposited in CAS (8 holotype: CASENT 9029653, PBI _ OON 03234 View Materials ; ♀ allotype CASENT 9029669, PBI _ OON 03376 View Materials )   .

ETYMOLOGY: The species is named after the type locality.

DIAGNOSIS: The male differs from others in the group by the large bulb-cymbium (about 0.7 patella-tibia length; but about 0.5 in other species) and the presence of a shallow dorsal groove proximally on the bulb (fig. 482). The female differs in having a smaller dorsal scute (fig. 279) and the epigynal region weakly sclerotized and the receptaculum short and relatively straight (W/L about 0.25) (figs. 296, 297).

MALE (PBI_ OON 03234 View Materials ). Total length 1.53, carapace length 0.74, width 0.62. CEPHALOTHORAX: Carapace (figs. 269, 270, 274) as in M. ranomafana   . Eyes as in M. ranomafana   , but slightly smaller; eye area about 0.5 carapace width, length subequal to clypeus length (figs. 269, 273). Sternum (figs. 272, 472) as in M. ranomafana   . ABDO- MEN (figs. 269, 271, 275–277) as in M. valterova   , but with slightly smaller book lung covers. GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 271). Palp with bulb-cymbium large, about 0.7 patellatibia length; bulb with dorsal depression (figs. 288–295, 482, map 3).

FEMALE (PBI_ OON 03376 View Materials ). Total length 1.82 (1.80–1.82), carapace length 0.78, width 0.68 (0.66–0.68), N 5 2. CEPHALOTHO- RAX (figs. 278–280, 283, 284): Clypeus sloping forward in lateral view, shorter than eye row length (fig. 280). Eye area about 1.3 clypeus length (figs. 280, 283). ABDOMEN (figs. 281, 285–287): Pedicel tube short (fig. 286), scutum dorsal extension slightly less than pedicel diameter (fig. 286). Dorsal scutum smaller than carapace, covering about 0.6 abdomen length and width (fig. 279). Postepigastric scutum weakly sclerotized. LEGS: patella plus tibia I longer than carapace (fig. 280). GENITALIA: Epigynal region weakly sclerotized, receptaculum short and relatively straight, W/L about 0.25 (figs. 296–300, map 4).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: Res. Andringitra, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, rainforest, sifting leaf litter, 23 Oct 1993 (B. Fisher, CASENT 9029654, PBI_ OON 03375 View Materials ), 1♀.

DISTRIBUTION: Known only from the type locality in Res. Andringitra, Madagascar.

Malagiella fisheri   , new species Figures 301–328 View Figs View Figs View Figs , 474 View Figs , 485 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifting leaf litter in rainforest, at 28.5 km S Midongy-Sud, Mount Papango, 23 ° 50.19S, 46 ° 57.89E, 940 m, P.N. Befotaka-Midongy, Fianarantsoa Province, Madagascar (13–15 Nov 2006, Fisher et al.), deposited in CAS (CASENT 9030777, PBI _ OON 03452 View Materials ).

ETYMOLOGY: This species is named after Brian Fisher, collector of this and several other species of Malagiella   , in honor of his impressive sampling program of the Malagasy fauna.

DIAGNOSIS: The male of this species differs from other Malagiella   in lacking sternal bristles (fig. 474) and in having a huge palpal patella (figs. 321–324, 485). The female differs from others in having a distinctly procurved posterior margin of the PES (figs. 327, 328) and a slightly curved receptaculum (W/L about 0.3).

MALE (PBI_ OON 03452 View Materials ): Total length 1.62, carapace length 0.82, width 0.69. CEPHALOTHORAX: Eye area about 0.5 carapace width; subequal to clypeus length (figs. 302, 303, 306). Sternum lacking stiff bristles found in other species (figs. 305, 474), median concavity absent. ABDOMEN (figs. 304, 308–310): as in M. valterova   . GENITALIA (figs. 321–326): Palp with patella greatly enlarged (fig. 485), about five times femur length, but less than four times in other species (figs. 481–484, 485).

FEMALE (PBI_ OON 03452 View Materials ): Total length 1.70 (1.70–1.86), carapace length 0.80 (0.78– 0.80), width 0.70 (0.68–0.70), N 5 2. CEPHALOTHORAX (figs. 311–313): Clypeus shorter than eye region length. Eye area about 1.8 clypeus length (fig. 313). ABDO- MEN (figs. 314, 318–320): Pedicel tube (fig. 337) shorter than in male. GENITA- LIA: Postepigastric scutum with posterior margin procurved, receptaculum only slightly curved, W/L about 0.3 (figs. 327, 328).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: P.N. Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23 ° 50.59S, 46 ° 57.59E, 1250 m, montane rainforest, sifting leaf litter, 17–19 Nov 2006 (B. Fisher et al., CASENT 9030776, PBI_ OON 03442 View Materials ), 1♀.

DISTRIBUTION: Known only from P.N. Befotaka-Midongy, Madagascar.

Malagiella andringitra   , new species

Figures 268–300 View Figs View Figs View Figs View Figs , 472 View Figs , 482 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype sifted from leaf litter in rainforest, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, Res. Andringitra, Fianarantsoa Province, Madagascar (23 Oct 1993, B. Fisher), deposited in CAS (8 holotype: CASENT 9029653, PBI _ OON 03234 View Materials ; ♀ allotype CASENT 9029669, PBI _ OON 03376 View Materials )   .

ETYMOLOGY: The species is named after the type locality.

DIAGNOSIS: The male differs from others in the group by the large bulb-cymbium (about 0.7 patella-tibia length; but about 0.5 in other species) and the presence of a shallow dorsal groove proximally on the bulb (fig. 482). The female differs in having a smaller dorsal scute (fig. 279) and the epigynal region weakly sclerotized and the receptaculum short and relatively straight (W/L about 0.25) (figs. 296, 297).

MALE (PBI_ OON 03234 View Materials ). Total length 1.53, carapace length 0.74, width 0.62. CEPHALOTHORAX: Carapace (figs. 269, 270, 274) as in M. ranomafana   . Eyes as in M. ranomafana   , but slightly smaller; eye area about 0.5 carapace width, length subequal to clypeus length (figs. 269, 273). Sternum (figs. 272, 472) as in M. ranomafana   . ABDO- MEN (figs. 269, 271, 275–277) as in M. valterova   , but with slightly smaller book lung covers. GENITALIA: Epigastric region with sperm pore small, rebordered (fig. 271). Palp with bulb-cymbium large, about 0.7 patellatibia length; bulb with dorsal depression (figs. 288–295, 482, map 3).

FEMALE (PBI_ OON 03376 View Materials ). Total length 1.82 (1.80–1.82), carapace length 0.78, width 0.68 (0.66–0.68), N 5 2. CEPHALOTHO- RAX (figs. 278–280, 283, 284): Clypeus sloping forward in lateral view, shorter than eye row length (fig. 280). Eye area about 1.3 clypeus length (figs. 280, 283). ABDOMEN (figs. 281, 285–287): Pedicel tube short (fig. 286), scutum dorsal extension slightly less than pedicel diameter (fig. 286). Dorsal scutum smaller than carapace, covering about 0.6 abdomen length and width (fig. 279). Postepigastric scutum weakly sclerotized. LEGS: patella plus tibia I longer than carapace (fig. 280). GENITALIA: Epigynal region weakly sclerotized, receptaculum short and relatively straight, W/L about 0.25 (figs. 296–300, map 4).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: Res. Andringitra, 38 km S Ambalavo, 22 ° 129S, 46 ° 589E, 1680 m, rainforest, sifting leaf litter, 23 Oct 1993 (B. Fisher, CASENT 9029654, PBI_ OON 03375 View Materials ), 1♀.

DISTRIBUTION: Known only from the type locality in Res. Andringitra, Madagascar.

Malagiella fisheri   , new species Figures 301–328 View Figs View Figs View Figs , 474 View Figs , 485 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifting leaf litter in rainforest, at 28.5 km S Midongy-Sud, Mount Papango, 23 ° 50.19S, 46 ° 57.89E, 940 m, P.N. Befotaka-Midongy, Fianarantsoa Province, Madagascar (13–15 Nov 2006, Fisher et al.), deposited in CAS (CASENT 9030777, PBI _ OON 03452 View Materials ).

ETYMOLOGY: This species is named after Brian Fisher, collector of this and several other species of Malagiella   , in honor of his impressive sampling program of the Malagasy fauna.

DIAGNOSIS: The male of this species differs from other Malagiella   in lacking sternal bristles (fig. 474) and in having a huge palpal patella (figs. 321–324, 485). The female differs from others in having a distinctly procurved posterior margin of the PES (figs. 327, 328) and a slightly curved receptaculum (W/L about 0.3).

MALE (PBI_ OON 03452 View Materials ): Total length 1.62, carapace length 0.82, width 0.69. CEPHALOTHORAX: Eye area about 0.5 carapace width; subequal to clypeus length (figs. 302, 303, 306). Sternum lacking stiff bristles found in other species (figs. 305, 474), median concavity absent. ABDOMEN (figs. 304, 308–310): as in M. valterova   . GENITALIA (figs. 321–326): Palp with patella greatly enlarged (fig. 485), about five times femur length, but less than four times in other species (figs. 481–484, 485).

FEMALE (PBI_ OON 03452 View Materials ): Total length 1.70 (1.70–1.86), carapace length 0.80 (0.78– 0.80), width 0.70 (0.68–0.70), N 5 2. CEPHALOTHORAX (figs. 311–313): Clypeus shorter than eye region length. Eye area about 1.8 clypeus length (fig. 313). ABDO- MEN (figs. 314, 318–320): Pedicel tube (fig. 337) shorter than in male. GENITA- LIA: Postepigastric scutum with posterior margin procurved, receptaculum only slightly curved, W/L about 0.3 (figs. 327, 328).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: P.N. Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23 ° 50.59S, 46 ° 57.59E, 1250 m, montane rainforest, sifting leaf litter, 17–19 Nov 2006 (B. Fisher et al., CASENT 9030776, PBI_ OON 03442 View Materials ), 1♀.

DISTRIBUTION: Known only from P.N. Befotaka-Midongy, Madagascar.

Malagiella fisheri   , new species Figures 301–328 View Figs View Figs View Figs , 474 View Figs , 485 View Figs , 487 View Fig ; maps 1–4; table 1

TYPE: Male holotype and female allotype from sifting leaf litter in rainforest, at 28.5 km S Midongy-Sud, Mount Papango, 23 ° 50.19S, 46 ° 57.89E, 940 m, P.N. Befotaka-Midongy, Fianarantsoa Province, Madagascar (13–15 Nov 2006, Fisher et al.), deposited in CAS (CASENT 9030777, PBI _ OON 03452 View Materials ).

ETYMOLOGY: This species is named after Brian Fisher, collector of this and several other species of Malagiella   , in honor of his impressive sampling program of the Malagasy fauna.

DIAGNOSIS: The male of this species differs from other Malagiella   in lacking sternal bristles (fig. 474) and in having a huge palpal patella (figs. 321–324, 485). The female differs from others in having a distinctly procurved posterior margin of the PES (figs. 327, 328) and a slightly curved receptaculum (W/L about 0.3).

MALE (PBI_ OON 03452 View Materials ): Total length 1.62, carapace length 0.82, width 0.69. CEPHALOTHORAX: Eye area about 0.5 carapace width; subequal to clypeus length (figs. 302, 303, 306). Sternum lacking stiff bristles found in other species (figs. 305, 474), median concavity absent. ABDOMEN (figs. 304, 308–310): as in M. valterova   . GENITALIA (figs. 321–326): Palp with patella greatly enlarged (fig. 485), about five times femur length, but less than four times in other species (figs. 481–484, 485).

FEMALE (PBI_ OON 03452 View Materials ): Total length 1.70 (1.70–1.86), carapace length 0.80 (0.78– 0.80), width 0.70 (0.68–0.70), N 5 2. CEPHALOTHORAX (figs. 311–313): Clypeus shorter than eye region length. Eye area about 1.8 clypeus length (fig. 313). ABDO- MEN (figs. 314, 318–320): Pedicel tube (fig. 337) shorter than in male. GENITA- LIA: Postepigastric scutum with posterior margin procurved, receptaculum only slightly curved, W/L about 0.3 (figs. 327, 328).

OTHER MATERIAL EXAMINED: MADA- GASCAR: Fianarantsoa Province: P.N. Befotaka-Midongy, Papango 27.7 km S Midongy-Sud, Mount Papango, 23 ° 50.59S, 46 ° 57.59E, 1250 m, montane rainforest, sifting leaf litter, 17–19 Nov 2006 (B. Fisher et al., CASENT 9030776, PBI_ OON 03442 View Materials ), 1♀.

DISTRIBUTION: Known only from P.N. Befotaka-Midongy, Madagascar.

CAS

California Academy of Sciences

MCZ

Museum of Comparative Zoology

USNM

Smithsonian Institution, National Museum of Natural History

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Oonopidae

Genus

Malagiella

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Oonopidae

Genus

Malagiella

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Oonopidae

Genus

Malagiella

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Oonopidae

Genus

Malagiella