Cryptocanthon brevisetosus Howden, 1973

Cook, Joyce, 2002, A Revision Of The Neotropical Genus Cryptocanthon Balthasar (Coleoptera: Scarabaeidae: Scarabaeinae), The Coleopterists Bulletin (mo 1) 56, pp. 3-96 : 27-31

9. Cryptocanthon brevisetosus Howden

( Figs. 52–56 )

Cryptocanthon brevisetosus Howden 1973:42

Material Examined. Seventy-four specimens (41 males, 33 females).

Type Material Examined. Holotype: ‘‘ MEX [ICO], 8mi S. / Simojovel , Chi [apa]s / VI-10-1969 / J.M. Campbell,’’ ‘‘ Berlese sample / #23’’ ( CNCI) male . Paratypes (4): same locality as holotype, ‘‘ VII-5-1969 / Bright & Campbell ’’ ( CMNC) 1 male ; Ocosingo / Chi [apa]s, Vi-1-2- / 1969, Bright / & Campbell ’’ ( CMNC) 2 males, 1 female .

Non-type Material Examined. GUATEMALA: ‘‘ Baja Verapaz, 4.3 km E. Purulha , 1,680 m, 29.VI–3.VII.1993, J. Ashe & R. Brooks, cloud forest, FIT’’

( CMNC) 1 male; ‘‘ Baja Verapaz , 8 km S. Purulha, 1,660 m, 19.V.1991, R. Anderson, pine/cloud for. litt., 91-1’’ ( CMNC) 4 males, 3 females; same data except ‘‘ 20.V.1991, 91-4,5’’ ( CMNC) 2 males, 3 females; same data except ‘‘ 25.V.1991, 91-22’’ ( CMNC) 2 males; same locality, ‘‘ 27.V.1991, H. & A. Howden, dung trap ’’ ( CMNC) 1 male; ‘‘ Baja Verapaz , 14.5 km S. Purulha, 1,600 m, 21.V.1991, R. Anderson, riparian bottomland oak forest litter, 91-7’’ ( CMNC) 3 males, 3 females; same data except ‘‘ 26.V.1991, 91- 27’’ ( CMNC) 1 female; ‘‘ Baja Verapaz , 4.5 km S. Purulha, 1,630 m, 21.V.1991, R. Anderson, cloud forest litter, 91-6’’ ( CMNC) 1 female; same data except ‘‘ 24.V.1991, 91-19’’ ( CMNC) 1 female, ( CMNC) 1 female; ‘‘ Baja Verapaz , 7.5 km S. Purulha, 1,630 m, 26.V.1991, R. Anderson, cloud forest litter, 91-26’’ ( CMNC) 3 males, 2 females; ‘‘ Baja Verapaz, 3.5 km S. Purulha , Ranchito El Quetzal, 1,700 m, 30.VI.1993, R. Anderson, cloud for. litter, 93-28D’’ ( CMNC) 2 males; same data except ‘‘ 1,770 m, 1.VII.1993, 93-31H’’ ( CMNC) 1 female; ‘‘ Baja Verapaz , 7 km SSW Purulha, 1,630 m, 30.XI.1991, leg R. Baranowski ’’ ( CMNC) 1 male, 2 females; ‘‘ Baja Verapaz , 17.5 km N. Salama (on #5), 1,650 m, 28.VI.1993, R. Anderson, oak forest litter, 93-23’’ ( CMNC) 3 males; ‘‘ Izabal, Cerro San Gil , 8 km N. Las Escobas, 800 m, 13.VI.1993, H. & A. Howden ’’ ( CMNC) 1 male, 1 female; same locality ‘‘ 11–14.VI.1993, B.D. Gill ’’ ( BDGC) 3 males, 5 females; MEXICO: same locality as holotype, ‘‘ VII-5-1969, Bright & Campbell ’’ ( CMNC) 1 male; ‘‘ Chi [apa]s, Ocosingo Rd. , 71 km S. Palenque, 5.VII.83, Rt. 195, 2,300 ft, S. & J. Peck, rainforest litter in ravine’’ ( CMNC) 1 male; ‘‘ Chiapas, 6 km S.W. Ocosingo, 1,400 m, 16.IX.1992, R.S. Anderson, 92-105, wet oak/ pine forest litter’’ ( CMNC) 1 male; same data except ‘‘ 22.IX.1991, 91–116’’ ( CMNC) 6 males, 6 females; ‘‘ Chiapas, 12.5 km N.W. Ocosingo, 1,400 m, 16.IX.1992, R.S. Anderson, 92-106 oak/pine forest litter’’ ( CMNC) 2 males, 2 females.

Diagnosis. This species is distinguished by the following combination of characters: vertical surface of clypeus not foveate; pronotal hypomeron carinate; inflexed portion of elytron with basal fovea; metathoracic wings greatly reduced; pygidium with strong median convexity, sharply declivous basally; paramere apices narrow, angulate ( Fig. 55 ).

Description. Male. Total length 3.0– 4.2 mm; greatest width 1.7–2.6 mm. Head. Clypeus not strongly produced anteriorly ( Fig. 53 ). Clypeal teeth moderately developed, widely separated, slightly reflexed. Clypeal emargination evenly rounded; vertical surface moderately broad, not foveate. Apical margin of clypeus lateral to teeth slightly convexly arcuate. Disc of clypeus flattened, slightly concave; coarsely, densely punctate, punctures separated by less than one diameter. Clypeogenal suture indicated by narrow carina. Gena not produced, anterior margin even with clypeal margin. Vertex and gena coarsely, densely punctate. Pronotum. Moderately convex medially, laterally explanate, shallow depressions laterally in posterior half; shallow median longitudinal depression in posterior half. Anterior angles nearly right-angled. Sides anterior to lateral angles nearly straight; evenly, slightly arcuate posterior to lateral angles. Posterior angles obtusely rounded, not incised. Pronotum densely, shallowly punctate; punctures smaller on disc, large and annulate laterally; setae short, inconspicuous. Pronotal hypomeron convex, with large shallow punctures; longitudinal carina present posterolaterally. Elytra. Transversely and longitudinally moderately convex; lateral carinae not strongly elevated. Each elytron dorsally with seven striae; each stria indicated by two clearly wavy lines. Elytral intervals flat; each interval with row of minute setose punctures on each side adjacent to striae; setae short, inconspicuous. Inflexed portion of elytron broad, evenly narrowed apically, with distinct basal fovea adjacent to epipleuron; relatively smooth with fine setose punctures; striae eight and nine absent; stria ten indicated by row of coarse punctures adjacent to margin of epipleuron. Epipleuron with scattered, moderate-sized punctures in basal half. Wings. Metathoracic wings reduced to elongate knobs. Venter.

54) aedeagus, lateral; 55) parameres, dorsal. Scale line 5 0.5 mm.

Prosternum finely punctate. Mesosternum finely punctate medially, with shallow annulate punctures laterally. Meso-metasternal suture slightly roundly angulate medially. Metasternum slightly convex medially; minutely punctate anteromedially; posteriorly and laterally with large, shallow, annulate punctures. Legs. Profemur densely punctate ventrally. Protibia slightly broadened medially in apical half; not notched before apex; apex strongly lobed medially ( Fig. 52 ). Mesofemur and metafemur moderately finely punctate; some punctures elongate. Metatrochanter unmodified. Metatibia evenly curved, crenulate on inner margin, with apical swelling on inner side. Abdomen. Apical sternite evenly convex, densely punctate; punctures large, shallow, annulate. Pygidium with strong median convexity sharply declivous basally; with short longitudinal carina medially at apex of convexity; short, broad sulcus at base; punctures large, dense. Male genitalia. Parameres ( Figs. 54–55 ) with apices narrow, angulate, not setose.

Variation. Female. Total length 3.6–4.0 mm; greatest width 2.2–2.5 mm. Clypeal teeth more strongly developed. Tibiae unmodified.

Both sexes vary in the length of the carina on the posterior part of the pronotal hypomeron.

Bionomics. This species has been taken in flight intercept traps in cloud forest, in dung traps, and sifted from riparian bottomland oak forest litter, wet oak/pine forest litter and cloud forest litter at elevations between 800 m and 1,770 m. It has been collected during the months of May–July, September and November.

Geographic Distribution ( Fig. 56). This species occurs in Guatemala and the state of Chiapas in Mexico.

Figures

Figs. 52–55. Cryptocanthon brevisetosus. 52) left protibia, male; 53) clypeal outline;