Anepholcia brechlini Behounek & Kononenko

Behounek, Gottfried & Kononenko, Vladimir, 2011, A revision of the genus Anepholcia Prout, 1924 with description of three new species from South East Asia (Lepidoptera, Noctuidae). Revision of Pantheinae, Contribution IV, Zootaxa 3097, pp. 20-34 : 22-23

Anepholcia brechlini Behounek & Kononenko , sp. n.

( Figs. 7, 8 , 26, 34, 42)

Material examined. Holotype: male PHILIPPINES, PALAWAN, Mt. Magosaw, Mainit Brook's Point, 600–900 m, Bal leg., ex coll. Brechlin R., 3–6.x.1996, in coll. GB ( ZSM), genitalia slide 7322m, HT. Paratype. 1 female, PHILIPPINES, NEGROS, Mt. Kanlaon, W-Route via Mambucal, 600–1000 m, primary forest, Brechlin R. leg., vii.1998, in coll. GB ( ZSM), genitalia slide 7336f. Holotype and paratype in GB, to be deposited in ZSM.

Diagnosis. The new species comprises an allopatric sister-species pair with A. pygaria . Externally it differs from A. pygaria by somewhat smaller size, more strongly expressed brown orbicular and reniform, less expressed white suffusion in terminal part of forewing, more smooth antemedial, submedial, subterminal and terminal lines and more developed terminal band on hindwing with reduced yellow anal patch and yellow terminal line. Female differs by having of filiform antennae. In male genitalia it differs by smaller size of genitalia, shorter and more rounded valva, shorter, but wider uncus, extended medially; smaller central sclerit of valva; about half shorter extension of sacculus and presence of quadrangular clavus. Aedeagus similar to A. pygaria , but pouches of vesica somewhat larger. In female genitalia antrum short, weakly sclerotised, ribbed; ductus and corpus bursae shorter than in A. pygaria .

Description. Adult ( Figs. 7, 8 ). Wingspan: male 51 mm, female 55 mm. Head whitish, patagia pale ochrebrown, tegulae ochre-brown, bordered with white; thorax and abdomen reddish-brown, abdominal crest presents. Antennae of male bipectinate, with moderate pectinations to beyond middle, shortening gradually to serrations, filiform in apical third; those of female filiform. Forewing glossy, ground colour dark brown with ochre-reddish, darker in basal and subbasal fields, suffused with whitish scales; basal line marked by white streak on costal margin; antemedial line angled outward on M, slightly curved inward and angled outward on A; medial shadow, distinct, waved; orbicular dark, distinct in male or formed by bordering line in female; reniform distinct in male, traceable in female, medial shadow distinct and waved in male, diffused in female; postmedial line distinct, dentate in male, diffused in female; veins behind postmedial lines suffused with blackish; tornal mark white with brown, well expressed in both sexes; white suffusion behind reniform in costal area, less expressed than in A. pygaria in male, more diffused in female; postmedial line waved, diffused; subterminal line diffused, but traceable in male, more distinct in female; terminal line white, less dentate than in A. pygaria , forming row of semilunar marks along termen in male, diffused in female; fringes brownish-grey or ochre-brown. Hindwing bright-yellow, with broad terminal band; terminal line thin, less expressed than in A. pygaria , line extending along wing and forming yellow anal angle smaller than in A. pygaria ; fringes brown-grey. Male genitalia (Fig. 26). Uncus shorter and broader than in A. pygaria , extended medially, somewhat constricted subapically, with concave apex; tegumen with wide lobes, shorter than vinculum; juxta triangular, shield-like, with central comb; valva shorter and more rounded than in A. pygaria ; costa and ventral margin of valva heavily sclerotised, cucullus rounded, covered with strong setae; central part of valva with small sclerotised plate; sacculus narrow, with quadrangular clavus basally and strong horn-like acute extension curved dorsally, shorter than in A. pygaria , reaching mid of valva. Aedeagus rather wide, strong, carina with numerous scobinate cornuti; vesica projecting dorsally, extended medially, with two pouch-like subbasal diverticula larger than in A. pygaria and small subapical diverticulum. Female genitalia ( Fig. 34 ). Papillae anales wide, quadrangular; apophyses posteriores and anteriores longer than in A. pygaria , equal in length; antrum shorter and less sclerotised compared with A. pygaria , ribbed; ductus bursae very short, membranous, corpus bursae sacculate, with small appendix bursae anteriorly.

Etymology. The species is dedicated to the German lepidopterist R. Brechlin who explored the fauna of Philippines and South East Asia.

Distribution (Map 42). Philippines: Palawan, Negros. Moths have been collected in July and October.

Figures

FIGURES 1 – 8. Anepholcia spp., adults. 1 — A. pygaria, female, holotype, Sumatra (BMNH); 2 — A. pygaria, male, Sumatra (BMNH); 3 — A. pygaria, male, North Thailand (ZSM); 4 — A. pygaria, female, Sumatra (GB); 5 — A. pygaria, male, Sumatra (GB); 6 — A. pygaria, female, Sumatra (GB); 7 — A. brechlini, male, holotype, Philippines, Palawan (GB); 8 — A. brechlini female, paratype, Philippines, Negros (GB).

FIGURES 33 – 37. Anepholcia spp., female genitalia. 33 — A. pygaria, female, Sumatra, genit. slide Z 4165 GB (ZSM); 34 — A. brechlini, paratype, Negros, genit. slide 7336 GB (GB); 35 — A. talboti, Sumatra, genit. slide Z 4193 GB (ZSM); 36 — A. kobesi, paratype, Sumatra, genit. slide Z 4167 GB (ZSM); 37 — A. philippina, paratype, Philippines, Mindanao, genit. slide 7337 GB.