Kempynus Navás, 1912

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 30-31

Kempynus Navás, 1912

( Figs 13–16 )

(= Kalosmylus Krüger, 1913a . Type species: Osmylus incisus McLachlan, 1873: 112 )

(= Osmylinus Banks, 1913 . Type species: Osmylus longipennis Walker, 1853: 235 )

(= Clydosmylus New, 1983a . Type species: Clydosmylus montanus New, 1983a: 409 ) syn. nov.

Type species. Osmylus incisus McLachlan, 1873: 112 (by original designation).

Diagnosis. Ocelli present; wings elongate, relatively large, slightly to strongly falcate, wing apices acute or rounded; FW subcostal veinlets often forked, usually strongly sinuous, interlinking crossveins rarely present; both wings with RP with ca. 11–20 branches, crossveins irregular, outer gradate series distinct, elongate and sinuous; FW M fork near origin of RP1 or RP2 RP; MP branching at or beyond of the mid-length of wing, sometimes with posterior branch partially fused to CuA; HW MA and MP relatively divergent, crossveins slightly sinuous; FW CuA with few branches, HW CuA elongate pectinately branched; CuP in both wings greatly elongate, pectinate branched; FW A1 and A2 elongate pectinate branched.

Comments. The largest genus in the subfamily, with species displaying a distinct austral distribution in Australia, New Zealand, Chile and Argentina. The typically falcate and strongly patterned wings, frequently sinuous crossveins throughout is typical for Kempynus . New (1983a) described the monotypic genus Clydosmylus New based on a large, sexually dimorphic species from eastern Australia. In their phylogenetic analyses Winterton et al. (2017) suggested that the differentiation of Clydosmylus from Kempynus is not supported and that Clydosmylus should be synonymised with Kempynus , which is done herewith. Lacewings with larger wings often develop additional vein branching and crossveins associated as a structural requirement to support the larger wing membrane area. The main features separating Clydosmylus from Kempynus described by New (1983a) include the large wing size, more reticulate venation and more basal forking of vein MP in the FW; all these features could easily be ascribed to the larger wing area. The distinctive New Zealand Kempynus incisus ( Fig. 14 ) was noted as also possibly being present in Australia by New (1983a) based on two specimens (lacking genitalia) collected in Queensland. No additional specimens have been collected so this record is yet to be confirmed.

Included species.

K. acutus New, 1986b ( Australia)

K. citrinus ( McLachlan, 1873) ( New Zealand)

K. crenatus Adams, 1971 ( Argentina; Chile)

K. digoniostigma Oswald, 1994 ( Argentina; Chile)

K. falcatus Navás, 1912 ( Argentina; Chile)

K. incisus ( McLachlan, 1863) ( Australia (?), New Zealand)

K. kimminsi New, 1983a ( Australia)

K. latiusculus ( McLachlan, 1894) ( New Zealand)

K. longipennis ( Walker, 1853) ( Australia)

K. maculatus New, 1983a ( Australia)

K. millgrovensis New, 1983a ( Australia)

K. montanus ( New, 1983a) comb. nov. ( Australia)

K. striatus New, 1983a ( Australia)

K. thecatus New, 1983a ( Australia)

K. tjederi Oswald, 1994 ( Chile)

Figures

FIGURE 13. Kempyninae. Kempynus incisus (McLachlan), dorsal view.

FIGURE 14. Kempyninae. Kempynus incisus (McLachlan), courtship behaviour (photo: Gil Wizen).

FIGURE 15. Kempyninae. Kempynus acutus New, dorsal view.

FIGURE 16. Kempyninae. Kempynus incisus (McLachlan), wing venation (trichosors and vestiture omitted).