Allomedmassa day, Dankittipakul & Singtripop, 2014

Dankittipakul, Pakawin & Singtripop, Tippawan, 2014, Allomedmassa, a new spider genus from evergreen forests of Southeast Asia (Araneae: Corinnidae), Revue suisse de Zoologie 121 (1), pp. 15-31 : 21-24

Allomedmassa day sp. nov. Figs 7-8, 13-14, 20-25

HOLOTYPE: 3, Thailand, Loei Province, Phu Ruea National Park , evergreen forests surrounding park head office, 900 m, pitfall trap, 9-10 August 2006, leg. P. Dankittipakul (MHNG, PDC-502).

PARATYPES: Nakhon Ratchasima Province, Khao Yai National Park, 770 m, 15 October 2008, leg. S. Pimpisalee, 23 (MHNG, PDC-541). – Khao Yai National Park , evergreen hill forest at 1000 m along a trail to view point, 10 August 2006, leg. P. Dankittipakul, 2♀ (MHNG, PDC-502) . – Chiang Mai Province, Chom Thong District, Doi Inthanon National Park, Doi Inthanon , 1650-1700 m, evergreen hill forest, sifting, 11 January 2006, leg. P. Dankittipakul, 13, 1♀ (MHNG, PDC-258). – Doi Inthanon, 1650-1700 m, evergreen hill forest, pitfall trap, 1-25 August 2002, leg. P. Dankittipakul, 13, 2♀ (MHNG, PDC-278) .

DIAGNOSIS: Males of A. day sp. nov. can be recognized by a bifurcated RTA (Fig. 22), a small, triangular prolateral tubercle on the palpal tibia (Fig. 21), and a hook-shaped embolus with broad base (Fig. 20). Females can be recognized by greatly enlarged copulatory orifices fusing anteriorly (Figs 13, 24), and by tubular spermathecae (Figs 14, 25). Allomedmassa day sp. nov. can be distinguished from A. mae sp. nov. by the presence of ten and eight pairs (instead of four pairs) of reduced ventral spines on tibiae of leg I and II, respectively.

ETYMOLOGY: The first author wishes to dedicate this new species to his father.

Originally Hainan Chinese: day = father; invariable noun.

DESCRIPTION OF MALE (HOLOTYPE): Total length 7.7; prosoma 3.3 long, 2.8 wide; opisthosoma 4.4 long, 1.9 wide. Eye sizes and interdistances: AME 1.0, ALE 0.5, PME = PLE 0.7, AME-AME 0.9, AME-ALE 0.5, PME-PME 1.0, PME-PLE 1.5. Leg formula 4123. Leg measurements: Leg I 13.3 (3.6, 4.8, 2.7, 2.2), leg II 12.1 (3.3, 4.3, 2.7, 1.8), leg III 10.8 (3.0, 3.5, 2.7, 1.6), leg IV 14.0 (3.8, 4.6, 4.1, 1.5). Spination: Leg I: Fe, d-11, p-1; Ti, v-2222222222; Mt, v-22; leg II: Fe, d-11, p-1; Ti, v-22222222, Mt, v-22; leg III: Fe, d-11, p-11; Ti, p-11, v-12; Mt, v-22; leg IV: Fe, d-11; Ti, p-1, v-11, r-1; Mt, p-11, v-11, r-1.

Coloration and pattern (Fig. 7). Carapace and chelicerae dark brown; labium and palpal coxae brown, distal area slightly paler; sternum brown, margin distinctly darker; legs yellow, except anterior femora dark brown; femora and patellae with dark FIGS 9-14

Allomedmassa mae gen. & sp. nov. (9-12) and Allomedmassa day gen. & sp. nov. (13-14). (9) Left male palp, ventral view. (10) Ditto, prolateral view. (11) Ditto, retrolateral view. (12, 14) Internal genitalia, dorsal view. (13) Epigyne, ventral view. Abbreviation: P, prolateral tubercle on palpal tibia; RTA, retrolateral tibial apophysis of male palp.

greenish distal annulation; tibiae and metatarsi with subproximal and distal annulations. Opisthosoma elongate-ovoid, sparsely covered with black pubescence; anterior median pigmentation triangular, with blunt apex, occupying slightly less than half of opisthosoma length, its margin not clearly outlined; dorsum gray, medially with paired pale patches, followed by seven transverse chevrons and pre-anal ring; venter pale, with dark median band running from epigastric furrow to spinnerets.

Palp (Figs 20-23). Palpal tibia cylindrical, with slightly elevated apicoventral surface (Fig. 20); prolateral tubercle triangular, heavily sclerotized, its apex blunt (Fig. 21); RTA bifurcated in retrolateral view (Fig. 22); tegulum ovoid, slightly excavated meso-prolaterally, with sigmoid sperm duct running medially; embolus short, black, hook-shaped, apex acutely pointed, directed prolaterad.

DESCRIPTION OF FEMALE (PARATYPE): Total length 7.5; prosoma 3.2 long, 2.8 wide; opisthosoma 4.3 long, 2.0 wide. Eye sizes and interdistances: As in male. Leg formula 4123. Leg measurements: Leg I 12.9 (3.5, 4.7, 2.7, 2.0), leg II 12.0 (3.2, 4.2, 2.8, 1.8), leg III 10.7 (3.0, 3.4, 2.6, 1.7), leg IV 13.8 (3.7, 4.5, 4.1, 1.5). Spination: Leg FIGS 15-19

Allomedmassa mae gen. & sp. nov.; male holotype (15-17), female paratype (18-19). (15) Left male palp, ventral view. (16) Ditto, proventral view. (17) Ditto, retrolateral view. (18) Epigyne, ventral view. (19) Internal genitalia, dorsal view. Abbreviations: CO, copulatory orifice; E, embolus; FD, fertilization duct; ID, insemination duct; P, prolateral tubercle on palpal tibia; RTA, retrolateral tibial apophysis; S, spermatheca; SR, accessory sperm receptacle; ST, subtegulum; T, tegulum. Scale lines = 0.1 mm.

I: Fe, p-1, d-11; Ti, v-2222222222; Mt, v-22; leg II: Fe, p-1, d-11; Ti, v-22222222, Mt, v-22; leg III: Fe, p-11, d-111; Ti, p-11, v-12, r-11; Mt, p-11, r-11, v112; leg IV: Fe, d-111, p-1, r-1; Ti, p-11, v-112, r-11; Mt, p-11, v-112, r-11.

Coloration and pattern (Fig. 8). Carapace anteriorly dark brown, posteriorly yellowish brown; chelicerae, labium and palpal coxae brown, distal part slightly paler than proximal part; sternum yellowish brown; anterior legs dark brown, posterior legs yellow. Opisthosoma ovoid; anterior median pigmentation indistinct; dorsum dark gray, with pale folium; venter pale.

Genitalia (Figs 13-14, 24-25). Epigynal region convex, heavily sclerotized; copulatory orifices situated medially, greatly enlarged, anteriorly fused together; internal genitalia represented by elongated, strongly convoluted insemination ducts and slightly enlarged posterior spermathecae; fertilization ducts short, elliptical.

FIGS 20-25

Allomedmassa day gen. & sp. nov.; male holotype (20-23), female paratype (24-25). (20) Left male palp, ventral view. (21) Ditto, prolateral view. (22) Ditto, retrolateral view. (23) Palpal tibia, dorsal view. (24) Epigyne, ventral view. (25) Internal genitalia, dorsal view. Scale lines = 0.1 mm.

NATURAL HISTORY: Allomedmassa day sp. nov. inhabits evergreen hill forests at moderately high altitudes (770-1700 m). Females were collected by sifting decomposing leaves and organic litter.

DISTRIBUTION: Northern and northeastern Thailand.