Chlororithra missioniaria Oberthür, 1916, Oberthur, 1916

Hongxiang, Han, Hongmei, Li, Dayong & Xue, 2006, Revision of Chlororithra Butler, 1889 (Lepidoptera, Geometridae, Geometrinae), Zootaxa 1221, pp. 29-39: 36-38

publication ID 10.5281/zenodo.172575

persistent identifier

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scientific name

Chlororithra missioniaria Oberthür, 1916

stat. nov.

Chlororithra missioniaria Oberthür, 1916   stat. nov.

Figs. 7–8 View FIGURES 1 – 8

Chlororithra fea   var. missioniaria Oberthür, 1916   , Études Lépid. comp. 12: 116, pl. 389, fig. 3280, 3281. Lectotype ɗ ( ZFMK), [ China]: Tsekou.

Chlororithra fea   ab. missioniaria: Prout, 1935   , in Seitz, Macrolepid. World 4 (Suppl.): 10, pl. 2 f. Chlororithra fea missioniaria: Scoble, 1999   , Geometrid Moths of the World, a Catalogue 1: 149. [as synonym of fea   ].


Head: Antenna in male bipectinate to three­fouths, ciliate apically; with shaft pale brown and pectinations dark brown. Frons mostly whitish, slightly broader than that of C. fea   , middle width about 3 / 4 horizontal diameter of compound eye. Vertex white. Labial palpus with all the segments whitish on both outer and inner surfaces; segment I and II rough­scaled, segment III smooth and slightly elongate in female compared to male, exposed.

Thorax: Tegula mostly whitish. Dorsum white and yellow green on middle of metathorax; ventral side white. Length of forewing (base to apex) 16–17 mm in both male and female. Wing pattern slightly paler than C. fea   . Lines white, edged with pale green line or patch, forming patches on costa. Subbasal line wavy; antemedial line wavy, edged with pale green outward; postmedial line serrated, edged with pale green inward; antemedial and postmedial lines approaching each other towards inner margin; submarginal line fairly wavy, extending inwards along veins, connected with postmedial line, forming oval patches between these two lines, with an accompanying pale green patch on both sides; terminal line forming white round patch between veins. Discal spot pale green, slender and bent, extending to costa. Fringe white between veins, pale green on veins. Hindwing with streaks similar to forewing, black­brown patch absent. Underside whitish, much paler than that of C. fea   , streaks on upper side very weakly discernable.

Abdomen: Dorsum and upper lateral side yellow­green, white intersegments; venter and lower lateral side whitish. Male genitalia ( Figs. 15–18 View FIGURES 9 – 21 ) similar to those of C. fea   , but with the following differences: socii much broader, tapered apically; costa with basal process almost smooth, while more­or­less spinulose in C. fea   ; coremata very weak, while frequently well developed in C. fea   ; center of sternite 8 deeply and broadly depressed, inner margin straight and smooth, U­shaped, and lateral process smooth, sharp and tapered ( Fig. 20 View FIGURES 9 – 21 ). Female genitalia ( Fig. 19 View FIGURES 9 – 21 ) with apophyses posteriores long, about 8 times of apophyses anteriores in length. Sternite 7 strongly sclerotized. Ductus bursae long, about as long as corpus bursae, with posterior half weakly sclerotized, antrum absent. Corpus bursae oval, membranous; signum a rounded, sclerotized patch with a tooth­like process at middle.

Materials examined

CHINA: paralectotypes, Tsekou, P. Dubernard, 1895, Ex. Oberthür Coll. Brit. Mus. 1927 ­ 3, 8 males ( BMNH); lectotype, Thibet Tsekou, R.P. Dubernard (other label: Chlororithra fea Butler   , var. missioniaria Obthr.   ), 1 male ( ZFMK); paralectotypes, Tse Kou, R.P. Dubernard, 1895, 3 males ( ZFMK); Donglingshan, Beijing, China, 1300 m, 2001. VI. 23, ex. coll. Galsworthy, 1 female ( IZCAS); Beijing Mentougou Liyuanling, 2001. VI. 21, coll. Xue Dayong, 1 male ( IZCAS); Yunnan: Lijiang Yulongshang, 1962. VIII. 3, coll. Song Shimei, 1 female ( IZCAS), 2800–2850 m, 1984. VII. 16–20, coll. Liu Dajun, 1 male, 1 female ( IZCAS), coll. Chen Yixin, 1 female ( IZCAS); Donglingshan, Beijing, China, 1300 m, 2001. VI. 23, coll. Galsworthy, 2 males (AG Coll.).


This species was described by Oberthür as a variation of C. fea   on the basis of 19 male specimens from Tsekou collected by R. P. Dubernard from 1895 to 1900. Specimens of this series are represented by two distinct wing patterns: in one form there is a blackbrown apical patch on both the upper and under sides, and in the second form the blackbrown apical patch is absent. Prout (1933) suggested that the difference was “possibly seasonal.” After dissecting a long series of both forms, it was revealed that C. fea   and C. missioniaria   have quite different male and female genitalia. Seven syntypes of missioniaria   with a black­brown patch have the same genitalia as C. fea   and are transferred to that species.

There are two main features of variation in the male genitalia — the shape of the apex of tha valva and the number of subapical teeth on the gnathos. The shape of the apex of the valva is rounded in specimens from Beijing and Tsekou, and longer and more slender in specimens from Lijiang. The gnathos usually bears a single subapical tooth, but in one specimen from Lijiang, there are three, possibly representing be a mutation.

Prout (1935) pointed out that “Oberthür united the whole series under the one name, but in order to conserve the name we have assumed his first figure to be the type.” Here, we designate the specimen that Oberthür figured as 3280 as the lectotype.


China (Beijing, Yunnan (Tsekou [Deqin], Lijiang)).


Zoologisches Forschungsmuseum Alexander Koenig


Institute of Zoology, Chinese Academy of Sciences














Chlororithra missioniaria Oberthür, 1916

Hongxiang, Han, Hongmei, Li, Dayong & Xue 2006

Chlororithra fea missioniaria:

Scoble 1999


Prout 1935

var. missioniaria Oberthür, 1916

Oberthur 1916