Rhamphomyia, Meigen, 1822

Sinclair, Bradley J., Vajda, Élodie A., Saigusa, Toyohei, Shamshev, Igor V. & Wheeler, Terry A., 2019, Rhamphomyia Meigen of the Canadian Arctic Archipelago, Greenland and Iceland (Diptera: Empididae), Zootaxa 4670 (1), pp. 1-94 : 6-7

publication ID

https://doi.org/ 10.11646/zootaxa.4670.1.1

publication LSID

urn:lsid:zoobank.org:pub:5D56C44D-BBAE-4CE7-9184-8A6DDC8009A4

DOI

https://doi.org/10.5281/zenodo.3796920

persistent identifier

https://treatment.plazi.org/id/03C52A7B-EB19-FF90-FF13-FA0398E0F818

treatment provided by

Plazi

scientific name

Rhamphomyia
status

 

Key to male Rhamphomyia of the Canadian Arctic Islands, Greenland and Iceland

1 Prosternum clothed entirely with white to brownish silky, hair-like setae ( Fig. 3D View FIGURE 3 ); sternite 6 with cluster of strong golden setae; sternite 7 with pair of horn-like projections ventrally ( Fig. 3C View FIGURE 3 ).................... R. (Ctenempis) albopilosa Coquillett

- Prosternum bare; sternite 6 without row of golden setae; sternite 7 unmodified or modified different than above.......... 2

2 Cerci directed anteriorly ( Figs 6A View FIGURE 6 , 9C View FIGURE 9 , 12B View FIGURE 12 , 15B View FIGURE 15 ) (subgenus Dasyrhamphomyia )................................... 3

- Cerci directed posteriorly............................................................................... 6

3 Thorax short haired (setae shorter than gaps between rows) with acrostichals and dorsocentrals 1–2 serial, with brown vittae beneath rows; cerci with dark apex, with left cercus overlapping right at tip ( Figs 12B, C View FIGURE 12 ); hind tibia with anteroventral row of dark, stout, spine-like setae shorter than half tibial width.................................. R. (D.) leptidiformis Frey

- Thorax long haired (setae longer than gaps between rows) with acrostichals and dorsocentrals 2–4 serial, without brown vittae beneath rows; cerci not darkened at apex, with right cercus overlapping left at tip ( Figs 9D View FIGURE 9 , 15C View FIGURE 15 ); hind tibia without anteroventral row of dark, stout, spine-like setae..................................................................... 4

4 Cerci pale, elongate, extending to tergite 3, overlapping subapically in addition to apical overlap ( Figs 15B, C View FIGURE 15 )........................................................................................... R. (D.) nigrita Zetterstedt

- Cerci brownish to black, shorter, extending to tergite 5, only overlapping apically ( Figs 6A View FIGURE 6 , 9C View FIGURE 9 )....................... 5

5 Dichoptic; abdomen with setae longer than length of abdominal tergites; cerci blackish ( Fig. 6A View FIGURE 6 ); hind tibia and femur with pronounced fine, pale ventral pubescence ( Fig. 6C View FIGURE 6 ).................................... R. (D.) erinacioides Malloch

- Holoptic; abdomen with setae shorter than length of abdominal tergites; cerci pale brown ( Fig. 9C View FIGURE 9 ); hind tibia and femur with long setae beneath ( Fig. 9A View FIGURE 9 )...................................................... R. (D.) hovgaardii Holmgren

6 Wing without dm-m crossvein (cell dm open) ( Fig. 62C View FIGURE 62 ); scutum with shiny vittae................................................................................................. R. (Pararhamphomyia) diversipennis Becker

- Wing with dm-m crossvein (cell dm closed), sometimes dm-m crossvein or M 2 base absent in R. (P.) omissinervis Becker ; scutum without shiny vittae............................................................................. 7

7 Cercus with distinctive finger-like projection dorsally near base ( Figs 53B View FIGURE 53 , 60A View FIGURE 60 )................................... 8

- Upper margins of cercus without finger-like projections....................................................... 9

8 Large species (wing length more than 5 mm); phallus curvature occurring before folding into epandrium; cercal finger-like projection stout ( Fig. 53B View FIGURE 53 ); mid tibia with at least 3 preapical anteroventral and posteroventral setae dark, stout, spine-like..................................................................................... R. hirtula Zetterstedt

- Small species (wing length less than 4 mm); phallus curvature hidden within epandrium; cercal finger-like projection slim, short, less than half cercal length ( Fig. 60A View FIGURE 60 ); mid tibia with 2 anterodorsals and 2 posterodorsals longer than twice tibial width................................................................................ R. setosa Coquillett

9 Tergite 8 long, subequal to length of epandrium ( Iceland only) ( Fig. 46B View FIGURE 46 ).................... R. (P.) simplex Zetterstedt

- Tergite 8 shorter than length of epandrium................................................................ 10

10 Subepandrial sclerite prolonged into pair of medioposterior lobes beneath cerci ( Figs 31C View FIGURE 31 , 28A View FIGURE 28 , 25C View FIGURE 25 , 34A View FIGURE 34 , 36C View FIGURE 36 , 38C View FIGURE 38 , 40A View FIGURE 40 ).. .................................................................................................. 11

- Cercus without pair of subepandrial lobes beneath.......................................................... 17

11 Apex of subepandrial lobe forked....................................................................... 12

- Apex of subepandrial lobe unforked..................................................................... 14

12 Phallus stout, looping dorsally above cercus and epandrium; apex of phallus expanded with ridge of teeth-like projections ( Figs 28A, B View FIGURE 28 ).............................................................................. R. (P.) helleni Frey

- Phallus slender, filamentous apically, extending slightly beyond epandrium; tip of phallus slender, tapered, lacking teeth-like projections......................................................................................... 13

13 Fore tarsomere 1 swollen, slightly broader than width of fore tibia at apex ( Fig. 31A View FIGURE 31 ); tergite 5 without stout posterolateral setae; subepandrial lobe with outer finger-like process lacking apical seta, longer than inner lobe ( Figs 31 View FIGURE 31 C-E)............................................................................................ R. (P.) hilariformis Frey

- Fore tarsomere 1 slender, narrower than width of fore tibia at apex; tergite 5 with several closely set stout posterolateral setae ( Fig. 38B View FIGURE 38 ); subepandrial lobe with slender, outer finger-like process bearing 1 very long, fine, black seta ( Figs 38C, D View FIGURE 38 ).................................................... R. (P.) lymaniana Sinclair, Vajda, Saigusa & Shamshev sp. nov.

14 Phallus not forming loops.............................................................................. 15

- Phallus forming loops................................................................................. 16

15 Hind tibia with some anterodorsal, posterodorsal and dorsal setae more than 2X tibial width ( Fig. 25B View FIGURE 25 ); apex of subepandrial lobe rounded ( Figs 25A, C View FIGURE 25 )............................ R. (P.) frigida Sinclair, Vajda, Saigusa & Shamshev sp. nov.

- Hind tibia with anterodorsal, posterodorsal and dorsal setae shorter than 2X tibial width ( Fig. 34B View FIGURE 34 ); apex of subepandrial lobe pointed ( Fig. 34A View FIGURE 34 )....................................................................... R. (P.) hoeli Frey

16 Hind tibia clavate ( Fig. 36D View FIGURE 36 ); hind tarsomere 1 shorter than length of tarsomeres 2–4 combined ( Fig. 36D View FIGURE 36 )............................................................................................ R. (P.) kjellmanii Holmgren

- Hind tibia not clavate ( Fig. 40B View FIGURE 40 ); hind tarsomere 1 longer than length of tarsomeres 2–4 combined ( Fig. 40B View FIGURE 40 )........................................................................................... R. (P.) omissinervis Becker

17 Phallus hidden within epandrium, leaving only base visible ( Fig. 17B View FIGURE 17 )................................................................................. R. (Eorhamphomyia) shewelli Sinclair, Vajda, Saigusa & Shamshev sp. nov.

- Phallus slim to hair-like, usually extending farther than length of epandrium, and often with curvatures................ 18

18 Phallus with distinct loop on apical half around apex of epandrium ( Figs 51C View FIGURE 51 , 57A View FIGURE 57 )............................... 19

- Phallus without loop, gradually curved................................................................... 20

19 Sternite 8 with stout, “horn-like” lateral projections ( Fig. 51D View FIGURE 51 ); phallus with loop forming acute angle ( Fig. 51C View FIGURE 51 ); hind tarsomere 1 swollen, wider than hind tibial width at apex ( Fig. 51B View FIGURE 51 )................................ R. herschelli Malloch

- Sternite 8 without projections; phallus with small in-ward U-shaped loop ( Fig. 57A View FIGURE 57 ); hind tarsomere 1 slender ( Fig. 56B View FIGURE 56 )........................................................................................ R. laevigata Loew

20 Midleg distinctly modified: femur strongly curved upwards; tibia curved inwards, with 3–5 dark, stout mid-posterodorsal setae longer than 2X tibial width; tarsomere 1 spindle-shaped, covered with dense, erect and moderately long setae ( Figs 42A, B View FIGURE 42 )................................................ R. (P.) petervajdai Sinclair, Vajda, Saigusa & Shamshev sp. nov.

- Midleg simple....................................................................................... 21

21 Apex of epandrium truncate ( Figs 44A, C View FIGURE 44 ); hind tibia with distinctive inward bend on apical part; hind tarsomere 1 spindleshaped, convex dorsally but straight ventrally ( Fig. 44B View FIGURE 44 )................................................................................................... R. (P.) septentrionalis Sinclair, Vajda, Saigusa & Shamshev sp. nov.

- Apex of epandrium rounded; hind tibia straight; hind tarsomere 1 clavate or uniformly slender....................... 22

22 Anepisternum shiny and scutum with 1–2 pairs of shiny vittae; hind tarsomere 1 clavate, increasing in size apically such that apical width broader than hind tibia ( Fig. 48B View FIGURE 48 ); cerci weakly constricted in middle, posterior half parallel to epandrium ( Fig. 48A View FIGURE 48 ).......................................................................... R. (P.) ursinella Melander

- Anepisternum pruinose and scutum with pruinescent vittae; hind tarsomere 1 evenly slender, nearly as broad as apical width of hind tibia, ( Fig. 25B View FIGURE 25 ); cerci strongly constricted in middle, posterior half strongly divergent from epandrium ( Fig. 25A View FIGURE 25 )....................................................................... R. (P.) filicauda Henriksen & Lundbeck

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Chironomidae

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