Clubiona conica, Dankittipakul & Singtripop, 2014

Dankittipakul, Pakawin & Singtripop, Tippawan, 2014, A new species-group of Clubiona Latreille, 1804 and descriptions of four new species from Borneo (Araneae, Clubionidae), Journal of Natural History (J. Nat. Hist.) 48 (31 - 32), pp. 1923-1936 : 1930-1933

Clubiona conica sp. nov.

( Figures 1D , 2A–C , 4A–E )

Diagnosis

The male of C. conica sp. nov. can be recognized by the long, sickle-shaped retrolateral apophysis on the palpal tibia, and by the very short, spiniform embolus ( Figures 1D , 4A–C ). The female is recognized by the thin-walled insemination ducts connected to pyriform spermathecae, and by the membranous bursae with distinct, elongated stalk ( Figures 2A , 4D,E ).

Type material

Holotype. ♂, Malaysia, Borneo, Sabah State, Kinabalu National Park, Poring Hot Spring , 500 m, 8 May 1987, D. Burckhardt & I. Löbl leg. (MHNG, 16H).

Paratype. One ♀, same data as for holotype (MHNG, 16H) .

Etymology

The specific epithet is derived from the Greek (κωΝΟϛ = conus = cone-shaped), alluding to a conical base of embolus.

Description

Male (holotype). Total length 5.8; prosoma 3.1 long, 2.8 wide; opisthosoma 2.7 long, 1.3 wide. Prosoma ovoid, pars cephalica broad, almost as wide as carapace width, anterior margin straight, in profile flat; integument smooth; fovea shallow, longitudinal; carapace orange brown, with distinctly brown anterior margin; chelicerae dark brown, with three promarginal and four retromarginal teeth; labium and gnathocoxae brown; sternum pale yellow, bordered with brown margin. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.22, PLE 0.20, AME–AME 0.10, AME–ALE 0.10, PME–PME 0.62, PME–PLE 0.30; MOQ: 0.40 long, 0.30 anterior width, 0.80 posterior width. Legs uniform pale yellow; scopulae on tarsi and metatarsi I–II distinct, in III–IV sparse. Leg formula 4213. Leg measurements: leg I 5.3 (1.6, 0.8, 1.4, 1.1, 0.4); leg II 6.6 (2.0, 0.8, 2.1, 1.2, 0.5); leg III 6.0 (1.5, 1.4, 1.2, 1.4, 0.5); leg IV 7.9 (2.2, 0.7, 2.0, 2.2, 0.8). Opisthosoma ovoid, pale, without colour marking; dorsal scutum lightly sclerotized, located on cardiac region, extending half length of opisthosoma, followed by pair of muscle apodemes.

Palp ( Figures 1D , 4A–C ). Palpal tibia short, approximately half length of cymbium; dorsal surface with blunt tubercle; ventro-prolateral surface with broad, triangular, sclerotized VTA; RTA sickle-shaped, greatly elongate, extending anteriorly then curving backwards, tapering towards sharply pointed apex; cymbium with apicodorsal excavation, covered with thick patch of hairs; small, blunt tubercle located on meso-retrolateral side of cymbium; tegulum oblong, apical region weakly sclerotized; sperm duct U-shaped; embolic base dark, conical, situated medially on apical surface of tegulum; embolus spiniform, minute, apex sharply pointed, directed retrolaterad, emerging on apex of embolic base.

Female (paratype). Total length 6.9; prosoma 3.5 long, 3.0 wide; opisthosoma 3.4 long, 1.8 wide. General appearance as in male, but entire body pale yellow, dorsal scutum on opisthosoma absent. Eye sizes and interdistances: AME 0.10, ALE 0.10, PME 0.20, PLE 0.20, AME–AME 0.10, AME–ALE 0.10, PME–PME 0.64, PME–PLE 0.32; MOQ: 0.46 long, 0.30 anterior width, 0.86 posterior width. Legs uniform yellow; scopulae on tarsi and metatarsi I–II distinct, in III–IV sparse. Leg formula 4213. Leg measurements: leg I 5.9 (1.8, 0.8, 1.6, 1.2, 0.5); leg II 7.3 (2.2, 0.8, 2.2, 1.4, 0.7); leg III 6.4 (1.6, 1.6, 1.2, 1.5, 0.5); leg IV 8.4 (2.4, 0.9, 2.0, 2.3, 0.8).

Genitalia ( Figures 2A–C , 4D,E ). Epigynal region weakly sclerotized, posteriorly with heavily sclerotized, oval depression situated near epigastric furrow; copulatory orifices circular, separated from one another by their diameter apart, located at centre of epigynal depression; insemination ducts thin-walled, ascending anteriorly, turning outwards and connected to dorsal surface of pyriform spermathecae; spermathecae ventrally with obliquely located falcate projection ( Figures 2B , 4D ), apically with digitiform glandular head ( Figures 2C , 4D,E ); fertilization ducts ribbon-shaped, distinctly elongate ( Figure 2B,C ); bursae membranous, with tubular stalk originating on dorsal surface of spermathecae.

Natural history

Clubiona conica sp. nov. is a lowland species. Type specimens were collected by sifting leaf and organic litter under dipterocarp trees.

Distribution

Known only from the type locality ( Kinabalu National Park , Borneo) .

Figures

Figure 1. Male palp, ventral view (A–E), prolateral view (F), dorsal view (G). Clubiona apiculata sp. nov. (A,F,G). Clubiona cylindriformis sp. nov. (B). Clubiona cultrata sp. nov. (C). Clubiona conica sp. nov. (D). Clubiona cf. hystrix (E).

Figure 2. (A,D) Internal female genitalia. (B) Falcate projection on spermatheca. (C,E) Glandular head on spermatheca. (F) Fertilization duct. (G) Carapace, dorsal view. (H) Right leg (first pair), prolateral view. (I) Left leg (first pair), prolateral view. (A–C) Clubiona conica sp. nov.; (D–I) Clubiona apiculata sp. nov.

Figure 4. Clubiona conica sp. nov.; male holotype (A–C); female paratype (D–E). Male palp; (A) ventral view, (B) retrolateral view, (C) dorsal view. (D) Epigyne, ventral view. (E) Internal genitalia, dorsal view. Scale bars: A–C, 0.1 mm; D,E, 0.25 mm.