Diplommatina shivapuriensis Budha & Backeljau, 2017

Budha, Prem B., Naggs, Fred & Backeljau, Thierry, 2017, The genus Diplommatina Benson, 1849 (Gastropoda: Caenogastropoda: Diplommatinidae) in Nepal, with the description of seven new species, European Journal of Taxonomy 337, pp. 1-30 : 19-20

Diplommatina shivapuriensis Budha & Backeljau sp. nov.

urn:lsid:zoobank.org:act:76F83F52-F806-48D2-A61F-57C5BA4A2E29

Figs 7C , 9L

Diagnosis

Measurements (n = 8): SH 2.5–3.0 mm, SW 1.2–1.3 mm, Wh 6.5–7.0; holotype: SH 2.9 mm, SW 1.4 mm, Wh 6.5. Dextral, penultimate and body whorl nearly equal, constriction at the middle of the parietal lip ( Fig. 9L ). Two parietalis: Prt1 long and well developed; Prt2 short and hardly visible ( Fig. 7C 3 ). One vertical palatalis along the constriction. One horizontal palatalis before the constriction, just above the umbilicus ( Fig. 9L ). Columellaris prominent. Protoconch smooth, Wh approx. 2.0, with minute pits evenly distributed, no radial ribs ( Fig. 7C 1 ). Teleoconch with radial ribs; about 8–10 ribs/ 0.5 mm on the penultimate whorl; 10–13 ribs/ 0.5 mm on the body whorl towards the aperture ( Fig. 7C 2 , C 4 ). Spiral striation present. Umbilicus closed. Aperture circular. Peristome slightly curved at AeP, two lips, closer at angular edge. Parietal lip of the peristome slightly raised. Both lips slightly reflected. Operculum paucispiral, corneous, thin, transparent, slightly concave, outer surface smooth, inner surface with weakly raised peripheral circular edge.

Etymology

The species name refers to the Shivapuri peak from where it was collected.

Type material

Holotype

NEPAL: Shivapuri-Nagarjun National Park , 200 m below Shivapuri peak above Baghdwar, 2500 m, 27.813284 N, 85.387712 E. 25 Apr. 2008, P.B. Budha leg. ( CDZMTU048 ).

Paratypes

NEPAL: 50 shells from the type locality, same data as holotype ( CDZMTU 049); 6 shells from Deurali, Phobrang, Langtang National Park, 3247 m, 28.1057 N, 85.3263 E, 6 Jun. 2007, P.B. Budha leg. ( CDZMTU 049b).

Remarks

Diplommatina shivapuriensis sp. nov. is similar to D. sherfaiensis Godwin-Austen, 1870 from Sherborsip, Jaintia Hill, India (RBINS Dautzenberg collection I.G. 10591, 6 shells). Yet, D. shivapuriensis sp. nov. has a thinner shell, with a constriction in the middle (in D. sherfaiensis the constriction is more towards the columellar edge). The horizontal palatalis is widely separated from the parietal wall in D. sherfaiensis (see image ID 1903.7.1.2254 at www.nhm.ac.uk), but closer to the parietal wall in D. shivapuriensis sp. nov.

Figures

Fig. 7. Diplommatina Benson, 1849 of Nepal. A. D. regularis Fulton, 1901 (CDZMTU045). B. D. salgharica sp. nov., holotype (CDZMTU046). C. D. shivapuriensis sp. nov., holotype (CDZMTU048). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig. 9. Palatal lamellae (palatalis) of species of Diplommatina. Dotted lines indicate the position of the constriction. Dark lines indicate the palatalis. A. D. abiesiana sp. nov. B. D. exserta Godwin- Austen, 1886. C. D. fistulata sp. nov. D. D. folliculus (L. Pfeiffer, 1846). E. D. godawariensis sp. nov. F. D. maipokhariensis sp. nov. G. D. miriensis Godwin-Austen, 1917. H. D. munipurensis Godwin- Austen, 1892. I. D. oviformis Fulton, 1901. J. D. regularis Fulton, 1901. K. D. salgharica sp. nov. L. D. shivapuriensis sp. nov. M. D. silvicola Godwin-Austen, 1886. N. D. sperata W.T. Blanford, 1862. O. D. syabrubesiensis sp. nov.

Fig. 3. Diplommatina Benson, 1849 of Nepal. A. D. abiesiana sp. nov., holotype (CDZMTU020). B. D. exserta Godwin-Austen, 1886 (CDZMTU024). C. D. fistulata sp. nov., holotype (CDZMTU025). The yellow arrow indicates the dorsal fold. Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.

Fig.1. Map of the localities and areas surveyed for Diplommatina spp. in Nepal.

Fig. 2. Shell terminology. A. D. regularis Fulton, 1901. B. D. folliculus (L. Pfeiffer, 1846). C. General outline of the lower part of a shell from Diplommatina species. Abbreviations: Prt = Parietalis (Prt1 = inner parietalis, Prt2 = outer parietalis); Pt = Palatalis; Hpt = Horizontal palatalis (Hpt1 = lower, Hpt2 = upper in case of two); Vpt = Vertical palatalis (Vpt1 = left side, Vpt2 = right side in case of two).

Fig. 4. Diplommatina Benson, 1849 of Nepal. A. D. folliculus (L. Pfeiffer, 1846) (CDZMTU028). B. D. godawariensis sp. nov., holotype (CDZMTU032). C. D. maipokhariensis sp. nov., holotype (CDZMTU034). Scale bar in images A2, A4, B2, B4, C2, C4 is 500 µm.