Tenedos medina, Martínez & Brescovit & Quijano, 2022

Martínez, Leonel, Brescovit, Antonio D. & Quijano, Luis G., 2022, Revealing the diversity of ant-eating spiders in Colombia I: morphology, distribution and taxonomy of the barronus group of the genus Tenedos O. Pickard-Cambridge, 1897 (Araneae: Zodariidae), Zootaxa 5130 (1), pp. 1-154 : 135-138

Tenedos medina sp. n.

Figs 96–99 ; 107 .

Type material. Holotype: COLOMBIA. Cundinamarca: Medina, Secondary forest , Pitfall trap, 524m [4°33′43.15″N, 73°21′6.95″W], C. Cortés leg., 6-18.VIII.1999 _ 5.I-2-II.2000, 1 ♂ ( MPUJ _ ENT 0061989 ) . Paratypes: same data as the holotype, 1 ♂ (ICN-Ar-12310) , 1 ♂ ( MPUJ _ ENT 0068184 ), 1 ♂ ( MPUJ _ ENT 0061990 ), 1 ♀ ( MPUJ _ ENT 0068177 ) .

Etymology. The specific name is a noun in apposition taken from the type locality.

Diagnosis. Males of Tenedos medina sp. n., resemble those of T. andes Jocqué & Baert, 2002 , and T. carlosprietoi sp. n., by the presence of basal retrolateral tibial apophysis (bRTA); rounded concave median apophysis (MA) (see Jocqué & Baert, 2002: 83, fig. 7A–B; figs 94A–D; 95A–B; 97A–D, 98A–B; 99A–F; 101A–D; 102A–B; 104A–D), but can be separated from those species by their median apophysis (MA) with retrolateral notch strongly accentuated; hook-shaped, tubular, long basal retrolateral tibial apophysis, as well by long, very wide, shovelshaped retrolateral tibial apophysis (RTA) ( Figs 97A–D , 98A–B ; 99A–F ). Females resemble those of Tenedos andes Jocqué & Baert, 2002 and T. carlosprietoi sp. n., by large medial field plate (MFP) ( Figs 94E–F ; 95C–D ; 97E–F , 98C–D ; 101E–F ; 102C–D ; 104E–F ), but are distinguished by short lateral borders (LB) very wide at base, delimiting a larger and rounded medial atrium (A) ( Figs 97E–F , 98C–D ).

Description. Male (Holotype, ICN-Ar-8023). Coloration ( Fig. 96A–B ): carapace dark red, dark in margins. Chelicerae with paturon brown, fangs brown-reddish. Endites light brown, white on anterior region. Labium and sternum brown. Legs: Coxae I–IV pale beige. Femora I–IV light yellow on base and median region, brown the rest of their extension. Patella I yellow, II–IV light brown. Tibiae I–IV yellow with several brown spots. Metatarsus I brown, II–IV yellow with dark spots on the base. Tarsi I–IV yellow. Abdomen: dorsally dark gray with ten white guanine spots organized as follows: two irregular spots very close to each other partially fused in medial region, anteriorly positioned; two spots smaller than previous ones, anteromedially positioned; a transversal and large spot, following by three spots, all medially positioned; two large spots medially joined, posteriorly positioned. Laterally dark gray with two thin and oblique stripes, extending to the posterior side. Ventrally light gray with two large anterior spots. Spinnerets light brown. Measurements: total length 7.41, carapace length 3.79, width 2.12, height 1.61. Clypeus height 0.71. Eye diameters and interdistances: AME 0.11, ALE 0.12, PME 0.10, PLE 0.15; AME–AME 0.28, AME–ALE 0.23, AME–PME 0.23, PME–PME 0.25, PME–PLE 0.39, ALE–PLE 0.25. Chelicerae 0.93 length. Sternum length 1.21, width 1.12. Legs: I—femur 1.84/ patella 0.91/ tibia 1.90/ metatarsus 1.61/ tarsus 0.97/ total 7.23; II—1.76/ 0.68/ 1.31/ 1.55/ 0.92/ 6.22; III—1.61/ 0.43/ 1.31/ 1.63/ 0.89/ 5.87; IV—2.32/ 0.79/ 1.74/ 2.12/ 1.51/ 8.48. Abdomen length 2.61. Legs spines pattern (only the differences from the general pattern): III—femur d0-0-1, p0-1-1, metatarsus p1d-1-2. Palp: retrolateral process of cymbium (RPC) long, wider at base with a rounded projection at anterior side; tegulum (T) large, rounded, almost as long as wide, retrolateral excavation very accentuated; subtegulum (St) large, longer than wide, globose; conductor (C) developed, wide with granulated cuticle; appendix (ApC) short, apically sharp; embolus (E) long, filiform towards apex; base of embolus (EB) approximately as long as two times as basal tegular membrane; basal tegular membrane (BTM) originated proximally on tegulum, proximally flattened, ending as short and sharp appendix; spermatic ducts (SD) S-shaped thin with both folds full open; ventral tibial apophysis short and quadrangular-shaped in ventral view; median apophysis (MA) large, rounded with strongly notch on external edge; retrolateral tibial apophysis (RTA) large, longer than palpal tibia, shovel-shaped; basal retrolateral tibial apophysis (bRTA) long, quadrangular, hook-shaped ( Figs 97A–D , 98A–B ; 99A–F ).

Female (Paratype, MPUJ_ENT 0068177). Coloration and abdominal pattern of spots as male ( Fig. 96C–D ). Measurements: total length 8.03, carapace length 3.41, width 2.17, height 1.84. Clypeus height 0.67. Eye diameters and interdistances: AME 0.08, ALE 0.12, PME 0.12, PLE 0.16; AME–AME 0.23, AME–ALE 0.30, AME–PME 0.23, PME–PME 0.28, PME–PLE 0.44, ALE–PLE 0.31. Chelicerae 1.42 length. Sternum length 1.14, width 1.15. Legs: I—femur 1.98/ patella 0.64/ tibia 1.58/ metatarsus 1.42/ tarsus 1.01/ total 6.63; II—1.69/ 0.81/ 1.26/ 1.28/ 0.81/ 5.85; III—1.57/ 0.71/ 0.98/ 1.37/ 0.74/ 5.37; IV—2.07/ 0.93/ 1.68/ 2.15/ 0.93/ 7.76. Abdomen length 4.09. Legs spines pattern (only the differences from the general pattern): I, II—tibia v1r-1r-2. III—femur d0-0-1d, p1d-0-0. Epigyne: lateral borders (LB) long, curved towards posteromedial region of the epigyne, forming large medial atrium (A); median field plate (MFP) very large, quadrangular-shaped; copulatory ducts (CD) very short, wide, almost undistinguished from spermathecae; seminal receptacles (SR) long, wide, curved towards median septum; spermathecae (S) large, irregulars, posteriorly positioned; fertilization ducts (FD) almost as long as spermathecae length ( Figs 97E–F , 98C–D ).

Variation. Males (n=4): total length: 6.92–7.43; carapace length: 3.51–3.81; femur I length: 1.80–1.86.

Distribution. Known from Cundinamarca department ( Fig. 107 ).

Figures

FIGURE 94. Tenedos andes Jocqué & Baert, 2002. A–B. Male palp (AMNH_IZC 00217585): A. Prolateral view. B. Retroventral view. C. Ventral view. D. Retrolateral view. E–F. Female epigyne (ICN-Ar-8021): E. Ventral view. F. Dorsal view. Scale bars: A–B: 200µm; C–D: 500µm; E–F: 100µm.

FIGURE 95. Tenedos andes Jocqué & Baert, 2002. A–B. Male palp (AMNH_IZC 00217585): A. Ventral view. B. Retrolateral view. C–D. Female epigyne (ICN-Ar-8021): C. Ventral view. D. Dorsal view. Scale bars: A–B: 200µm; C–D: 500µm. Abbreviations: A, atrium; ApC, appendix of the conductor; bRTA, basal retrolateral tibial apophysis; C, conductor; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; MFP, median field plate; RPC, retrolateral process of the cymbium; RTA, retrolateral tibial apophysis; S, spermathecae; SD, spermatic ducts; SR, seminal receptacles; T, tegulum; VTA, ventral tibial apophysis.

FIGURE 96. Tenedos medina sp. n. A–B. Male habitus (MPUJ_ENT 0061989): A. Dorsal view. B. Ventral view. C–D. Female habitus (MPUJ_ENT 0068177): C. Dorsal view. D. Ventral view. Scale bar: A–D: 2mm.

FIGURE 97. Tenedos medina sp. n. A–B. Male palp (MPUJ_ENT 0061989): A. Prolateral view. B. Retroventral view. C. Ventral view. D. Retrolateral view. E–F. Female epigyne (MPUJ_ENT 0068177): E. Ventral view. F. Dorsal view. Scale bars: A–D: 500µm; E–F: 100µm.

FIGURE 98. Tenedos medina sp. n. A–B. Male palp (MPUJ_ENT 0061989): A. Ventral view. B. Retrolateral view. C–D. Female epigyne (MPUJ_ENT 0068177): C. Ventral view. D. Dorsal view. Scale bars: A–B: 200µm; C–D: 500µm. Abbreviations: A, atrium; ApC, appendix of the conductor; bRTA, basal retrolateral tibial apophysis; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; MFP, median field plate; RTA, retrolateral tibial apophysis; SR, seminal receptacles; VTA, ventral tibial apophysis.

FIGURE 99. Tenedos medina sp. n. A–F. Male palp (MPUJ_ENT 0061989), SEM images:A. Ventral view (arrow indicates the bRTA). B. Retrolateral view. C. Median apophysis. D. Surface of the conductor showing the granulated cuticle. E. Retrolateral tibial apophysis. F. Retrolateral tibial apophysis and tegulum (the arrow indicates the retrolateral excavation of the tegulum). Abbreviations: bRTA, basal retrolateral tibial apophysis; EB, embolus base; C, conductor; E, embolus; MA, median apophysis; RPC, retrolateral process of the cymbium; RTA, retrolateral tibial apophysis; St, subtegulum; VTA, ventral tibial apophysis.

FIGURE 101. Tenedos carlosprietoi sp. n. A–B. Male palp (IAvH-I-605): A. Prolateral view. B. Retroventral view. C. Ventral view. D. Retrolateral view. E–F. Female epigyne (IAvH-I-3007): E. Ventral view. F. Dorsal view. Scale bars:A–D: 500µm; E–F: 100µm.

FIGURE 102. Tenedos carlosprietoi sp. n. A–B. Male palp (IAvH-I-605): A. Ventral view. B. Retrolateral view. C–D. Female epigyne (IAvH-I-3007): C. Ventral view. D. Dorsal view. Abbreviations: A, atrium; bRTA, basal retrolateral tibial apophysis; E, embolus; FD, fertilization ducts; LB, lateral borders; MA, median apophysis; MFP, median field plate; RTA, retrolateral tibial apophysis; S, spermathecae; SR, seminal receptacles.

FIGURE 104. Tenedos carlosprietoi sp. n. A–B. Male palp (IAvH-I-606): A. Prolateral view. B. Retroventral view. C. Ventral view. D. Retrolateral view. E–F. Female epigyne (IAvH-I-614): E. Ventral view. F. Dorsal view. Scale bars:A–D: 500µm; E–F: 100µm.

FIGURE 107. Distribution map of the species belonging to barronus species group in Colombia.