Umbrageocoris maai maai, Kóbor, 2019

Kóbor, Péter, 2019, Contributions to the knowledge of Umbrageocoris (Heteroptera: Lygaeoidea: Geocoridae), Zootaxa 4652 (2), pp. 384-390 : 386-388

Umbrageocoris maai maai new subspecies

( Figures 1B , 5 )

Material studied. Holotype: BRITISH N. BORNEO / Tenompok, 1460m / Jesselton, 48km / E., 17–18. X. 1958 / T. C. Maa / Collector / BISHOP (1 ♂, BPBM) . Paratypes: BORNEO: SARAWAK / Bau, Lake Area / 29. VIII. 1958 // T. C. Maa / Collector / BISHOP (1 ♀, BPBM) ; NORTH BORNEO / Ranau / 28–30. IX. 1958 // Crotalaria / L. W. Quate / Collector (1 ♀, BPBM) ; BRITISH N. BORNEO: / Ranau / 30. IX. –5. X. 1958 // T. C. Maa / Collector / BISHOP (1 ♀, BPBM) ; BRITISH N. BORNEO / Tenompok, 1460m / Jesselton , 48km / E., 17–18. X. 1958 / T. C. Maa / Collector / BISHOP (1 ♀, 1 ♂, BPBM) ; NORTH BORNEO / Tenompok, Mt. / Kinabalu , 3. XI. 58 // T. C. Maa / Collector / BISHOP (1 ♀, 1 ♂, BPBM) ; MALAYA: Pahang / Kuala Tahun / 12–14. XII. 1958 // T. C. Maa / Collector (1 ♀, BPBM) ; BRITISH N. BORNEO / Tenompok / 10–14. II. 1959 // T. C. Maa / Collector / BISHOP (1 ♀, BPBM) ; BRITISH N. BORNEO / Tawun, Quion Hill / 8–14. VII. 1962 / H. Holtmann / Malaise Trap / BISHOP (1 ♀, BPBM)

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Description. COLOURATION: Head ochraceous; base and posterior part of eyestalks with brownish markings of various extents. Eyes dark reddish to purplish, ocelli hyaline. Antennal segments I & IV ochraceous with at most apex brownish; segments II-III dark brownish to blackish. Labial segments uniformly ochraceous. Thorax. Pronotum ochraceous with dark brownish punctation and markings across pronotal callosities and posterior to them. Humeral angles with round brownish spots. Scutellum uniformly dark brownish. Hemelytra ochraceous with brownish punctures; oval brownish spot extending from middle of M-R to median part of membrane. Thorax later- ally and ventrally dark brownish with collar of prosternite, legs and peritreme of MTSEA ochraceous. Pleurites with irregular ochraceous spot at dorsal-posterior angles. Abdomen dark brownish with dorsal edge of connexiva and abdominal trichobothria ochraceous.

STRUCTURE (only differences from U. kondorosyi noted): Head. Median furrow at apical region of vertex present, but less expressed and shorter (compare Fig. 2C - D ). Ratio of antennal segments: 1: 2.33: 1.89: 2.56. Ratio of labial segments: 1: 0.71: 0.95: 1.1. Thorax. Pronotum with sparser, but coarser punctation as in U. kondorosyi . Surface anterior to callosities with more than one line of punctures; pronotal callosities only slightly bulging. Pronotum posterior to callosities narrowly impunctate at lateral margin, humeral angles and posterior margin. Pronotum median length to basal width: 1: 1.65. Scutellum with dense punctation, finer as on pronotum; trifurcate carina of scutellum with apices interrupted by punctures, but almost complete. Median length of scutellum to basal width: 1: 0.91. Greatest distance between cubital punctures somewhat wider than basal width of clavus. Study of genital structures revealed no major differences between U. maai and U. kondorosyi .

MEASUREMENTS [holotype (range of paratypes)]: total body length: 3.88 (3.65–3.78); head length: 0.58 (0.45–0.50); head width: 1.58 (1.50–1.53); ocellar distance: 0.45 (0.45–0.50); antennal segments: 0.23-0.53-0.43- 0.58 (0.18-0.23–0.5-0.53–0.38-0.40–0.55-0.58); labial segments: 0.53-0.38-0.50-0.58 (0.48-0.5–0.38-0.4 –0.45- 0.50–0.53-0.58); pronotum length: 0.88 (0.80–0.83); pronotum width: 1.45 (1.33–1.42); scutellum length: 0.88 (0.80–0.90); scutellum width: 0.80 (0.73–0.78).

Diagnosis. Dark brownish decorations of vertex, pronotum and corium are less extensive than in U. kondorosyi . Pronotum with denser punctation and relatively extensive impunctate “collar” anterior to callosities. Trifurcate carina of scutellum is interrupted by punctures at apices.

Etymology. The species is named in honour of Tsing C. Maa, dipterist of Bernice P. Bishop Museum for collecting long series of geocorine bugs, thereby contributing to the knowledge of these taxa distributed in New Guinea and the Indomalayan archipelago ( Fig. 5 ).

Distribution. According to the specimens studied the subspecies is distributed in North Borneo and the Malay Peninsula ( Fig. 5 ).

Figures

FIGURE 1. Habitus of Umbrageocoris species: A. U. kondorosyi Kóbor, 2019 (holotype, HNHM); B. U. maai maai new sub- species (holotype, BPBM); U. maai timorensis new subspecies (holotype, BPBM).

FIGURE 2. Head of Umbrageocoris species: A. vertex, U. kondorosyi; B. ventral view, U. kondorosyi; C. frontal view of head, U. kondorosyi; D. frontal view of head, U. maai maai. (abbreviations: at—antenniferous tubecles, cl—clypeus, lt—labial trough, lts—suture of labial trough, mf—median longitudinal furrow, mp—mandibular plates, os—ocular sulcus).

FIGURE 5. Distribution of Umbrageocoris maai Kóbor, 2019, subspecies.