Marilia Müller 1880

Morse, John C., 2017, Lannapsyche and Marilia species of China (Trichoptera: Odontoceridae), Zootaxa 4320 (1), pp. 81-99 : 87-88

Marilia Müller 1880

Oláh & Johanson (2010) summarized the diagnostic characters for separating Oriental species of this genus. We agree with most of their opinions, especially for the pattern of vertexal setal warts and some of the genital characters, such as the groove pattern on segment IX; the shape of the apicolateral corners and the lateromedian apical lobes of tergum IX, and the central ridge pattern on segment X. Additionally, for identifying Chinese species, the following diagnostic characters are considered to be very useful. In each male forewing (1) fork 2 is sessile or petiolate and (2) MA is confluent with R5 for a characteristic distance or not. (3) In each hind wing the Cu1 stem is aligned with Cu1+MP or not. In male genitalia, (4) the ratio of length to height of segment IX in lateral view and (5) the ratio of length to width of tergum IX in dorsal view are diagnostic; (6) the shape and size of the coxopodite and (7) the apical segment (= harpago) of an inferior appendage and (8) the degree of development of the basomesal setose bump on a coxopodite vary distinctively among species; (9) The shapes of the phallus and the phallotremal sclerite are diagnostic. Shapes of various features of the female genitalia are distinctive, including shapes of (10–11) segments IX and X, (12) ventral lobes and (13) sclerotized ridges of segments IX and X, (14) the subgenital plate, and (15) the spermathecal sclerite.

Mosely & Kimmins (1953) found the proportions of maxillary palp segments of Australian species to be variable. This occurs also within the Chinese species, and sometimes variation happens even on the right and left palps for a single specimen. For example, in the holotype of Marilia qinlingensis n. sp., the right maxillary palp proportions from the first segment to the fifth segment are 1(0.65mm)/0.96/1/1/1.1 with the fifth segment being the longest, while it is 0.92/1/1.1/1/0.88 on the left palp, with the third segment being the longest. Therefore, we did not use the maxillary palp formula as a diagnostic character in this study.

Based on our observation of female genitalia of two Chinese species, distinctive differences exist between species. Therefore, the description and illustration for female specimens of this genus may add important diagnostic characters for species recognition and, possibly, eventual phylogenetic interpretation.