Microhoria heracleana Kejval, 2020

Kejval, Zbyněk & Chandler, Donald S., 2020, Generic revision of the Microhoriini with new species and synonymies from the Palaearctic Region (Coleoptera: Anthicidae), Acta Entomologica Musei Nationalis Pragae 60 (1), pp. 95-154 : 138-139

Microhoria heracleana Kejval , sp. nov. ( Figs 105–107 , 154 )

Type locality. Greece, Crete, Heraklion Prefecture, NE of Matala, Komos (camping), 35°00′33.04″N 24°46′01.02″E, alt. 92 m.

Type material. HOLOTYPE :: ‘ GREECE, Crete Iraklion Pref. Komos (camping) NE Matala, 12.v.2005; 92 m N 35 00 33.04 E 024 46 01.02; P. Chvojka leg. [p]’ ( NMPC). PARATYPES: 1 ♀, same data as holotype ( NMPC); 5 3 ♀♀, ‘ CRETE m., 30.iv.2013 Kalamaki env., sandy coast, 0 m a.s.l., S. Benedikt lgt. [p]’ ( ZKDC, SBPC); 3 5 ♀♀, ‘ GRECIA, Creta Sud, Kalamaki, VII.2013, leg.A. Fancello’ ( ADBC); 3 1 ♀, ‘Greece-Crete Agios Pavlos 23.5.2007 lgt. O. Konvička [p]’ ( ZKDC); 2, ‘I. Jeniš lgt. Mirtos Kreta 3.6.1990 [p]’ ( ZKDC); 1, ‘Graecia-Creta M y r t o s 2.–4.vi.1990 ing. Kudrna lgt. [p]’ ( ZKDC); 1 ♀, ‘ CRETE: 12.6.1992 MATALA Iraklio pref. J. Batelka lgt. [p]’ ( ZKDC); 2 ♀♀, ‘ Creta Biró [p] // Herakleion 1906.VI. [p/h]’ ( NHMW); 4 2 ♀♀, ‘ Creta Biró [p] // Herakleion 1906.VI. [p/h] // Microhoria nectarina (Panz.) det. G. Uhmann 1983 [p/h]’ ( HNHM, DCDC); 1 ♀, same data, except: ‘Herakleion 1906. IV. [p]’ ( HNHM); 1, ‘ Kreta, Iraklion, 2.8.58 Eckerlein leg. [p]’ ( NHMW); 1, ‘ Kreta, Tympaki, 29.-31.7.58, Eckerlein lgt.’ ( NHMW); 1 ♀, ‘Gazi, Creta Mařan et Štěp. 1934. Coll. Bartoň[p]’ ( NMPC); 1, ‘ Kreta [h] // Microhoria nectarina (Panz.) det. G. Uhmann 1983 [p/h]’ ( HNHM); 2, ‘GRC, CRETE N, 4 km W Heraklion, Stomio, pláž 35°20′18″N, 25°04′15″E, 23.VI.2016 leg. David Frank [p; pláž = beach]’ ( ZKDC, DFPC).2 1 ♀, ‘ CRETA: Vai (spiaggia) 03.VII.2010 (su fiori di Timo) leg. M. Romano’ ( ADBC).

Description. Male (holotype). Body length 4.0 mm. Head brownish-black, mouthparts dark reddish; pronotum dark reddish, somewhat darkened antero-laterally; elytra yellowish, with extensive brownish-black markings ( Fig. 154 ); legs and antennae reddish, distal antennomeres somewhat darker.

Head 1.3 times as long as wide including eyes, moderately widely rounded posteriorly in dorsal view; tempora slightly narrowing posteriorly, posterior temporal angles indistinct. Eyes large, moderately convex. Surface glossy, distinctly punctate; punctures distinctly separate. Setation short, appressed to subdecumbent, with few short erect setae. Antennae only moderately enlarged over distal half; antennomeres X 1.2 times, XI nearly 2.4 times as long as wide.

Pronotum 1.1 times as long as wide, slightly narrower than head including eyes, nearly evenly rounded anteriorly, narrowed and slightly impressed postero-laterally in dorsal view; pronotal disc evenly and moderately convex. Surface glossy, distinctly punctate, similar to those on head. Setation as on head, with scattered, short erect setae, and some longer setae laterally at base.

Elytra 1.7 times as long as wide; humeri well-developed, postbasal impression indistinct; apices modified, channel of gland forming short tubular process at apical margin. Surface moderately glossy, densely punctate; punctation as on head, setation similar, subdecumbent, with sparsely scattered, short erect setae.

Legs simple.

Abdominal sternum VII simple. Tergum VII simple. Sternum VIII forming paired sclerites that are simply rounded postero-medially; tergum VIII simple. Aedeagus ( Figs 105–107 ): tegmen strongly hook-like and curved apically; endophallic armature with two pairs of robust spines (shorter curved, and longer straight).

Female. Identical with male for most external characters; elytral apices simple; sternum VII simple; tergum VII

subtriangular, shallowly excavate apically.

Variation. Body length (♀) 3.8–4.5 mm; head 1.3–1.4 times as long as wide.

Differential diagnosis. Microhoria heracleana sp. nov. belongs to the M. terminata species-group. It is undoubtedly very close to M. nectarina, differing by its more elongate head, and mainly by the narrow and strongly hooked apical part of the tegmen. For a habitus figure and male characters of the latter species see KEJVAL (2018).

Etymology. Named after Heracles, a divine hero in Greek mythology; adjective.

Distribution. Greece (Crete).

Figures

Figs 105–111. 105 – Microhoria heracleana sp. nov., aedeagus in lateral view; 106 – same, apex of tegmen in dorsal view; 107 – same, apex of everted endophallus, ventral view; 108 – M. impavida sp. nov., aedeagus in lateral view; 109 – M. inobscura (Pic, 1908), stat nov., aedeagus in lateral view; 110 – same, apical part in ventral view; 111 – M. kabulensis sp. nov., aedeagus in lateral (left) and ventral (right) view. Scale bars: 0.2 mm – A (Fig. 111), B (Figs 105–109).

Figs 151–163. 151–155 – Habitus: 151 – Microhoria bacillisternum sp. nov.; 152 – M. garavuti sp. nov.; 153 – M. gibbipennis sp. nov.; 154 – M. heracleana sp. nov., paratype, Stomio (ZKDC); 155 – M. truncatipennis (Pic, 1897) comb. nov. 156–161 – pronotum and base of head and elytra: 156 – M. angelinii (Degiovanni, 2012) comb. nov.; 157 – M. persica sp. nov., paratype (ZKDC); 158 – M. gibbipennis sp. nov.; 159 – M. truncatipennis; 160 – M. vosseleri (Pic, 1894); 161 – Microhoria sp., Spain, Jarama River env., near Madrid (ZFMK). 162 – M. depressa (LaFerté-Sénectère, 1849), Azerbaijan, Zarat (ZKDC), apex of female tergum VII; 163 – M. gibbipennis sp. nov., apex of female tergum VII.