Phortica (Ashima) efragmentata Toda, 2020

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 12

4) Phortica (Ashima) efragmentata Toda , sp. nov.

( Figs 2 , 3 D–G)

Diagnosis. Posterior piece of aedeagal median rod widely bifurcated ( Fig. 3F ); aedeagal free sclerite absent (ch.59- 0; Fig. 3F ).

Description. Supracervical setae 18–19. Dorsomedial, tentorial apodeme 3/4 as long as basal, parallel portion of dorsolateral, tentorial apodeme.Arista with arb = 8/4 ( Fig. 2A ). Palpus with hollow-shaped sense organ ( Fig. 2B ). Cibarial, medial sensilla 12–17 per side; posterior sensilla 6–7 per side ( Fig. 2C ). All tarsi entirely yellow ( Fig. 2G ); fore tarsomere I as long as 3 succeeding tarsomeres together; mid one slightly longer than rest tarsomeres together; hind one as long as rest together. Aedeagal sheath ventro-apically with a pair of strongly sclerotized postgonites much smaller than bifurcated pieces of aedeagal median rod ( Fig. 3F ).

Measurements (holotype, in mm): BL (straight distance from anterior edge of pedicel to tip of abdomen) = 3.65, ThL (distance from anterior notal margin to apex of scutellum) = 1.56, WL (distance from humeral cross vein to wing apex) = 2.46, WW (maximum wing width) = 1.08.

Indices (holotype, in ratio): FW/HW (frontal width/head width) = 0.44, ch/o (maximum width of gena/maximum diameter of eye) = 0.08, prorb (proclinate orbital seta length/posterior reclinate orbital seta length) = 1.78, rcorb (anterior reclinate orbital seta length /posterior reclinate orbital seta length) = 0.91, vb (subvibrissal seta length /vibrissa length) = 0.30, dcl (anterior dorsocentral seta length/posterior dorsocentral seta length) = (n/a), presctl (prescutellar seta length /posterior dorsocentral seta length) = 0.52, sctl (basal scutellar seta length /apical scutellar seta length) = (n/a), sterno (anterior katepisternal seta length/posterior katepisternal seta length) = (n/a), orbito (distance between proclinate and posterior reclinate orbital setae/distance between inner vertical and posterior reclinate orbital setae) = 2.08, dcp (distance between ipsilateral dorsocentral setae/distance between anterior dorsocentral setae) = 0.24, sctlp (distance between ipsilateral scutellar setae/distance between apical scutellar setae) = 1.20, C (2nd costal section between subcostal break and R 2+3 /3rd costal section between R 2+3 and R 4+5) = 2.35, 4c (3rd costal section between R 2+3 and R 4+5 /M 1 between r-m and dm-cu) = 1.50, 4v (M 1 between dm-cu and wing margin/M 1 between r-m and dm-cu) = 3.03, 5x (CuA 1 between dm-cu and wing margin/dm-cu between M 1 and CuA 1) = 1.00, ac (3rd costal section between R 2+3 and R 4+5 /distance between distal ends of R 4+5 and M 1) = 3.88, M (CuA 1 between dm-cu and wing margin/M 1 between r-m and dm-cu) = 0.66, C3F [length of heavy setation in 3rd costal section/(length of heavy setation in 3rd costal section+length of light setation in 3rd costal section)] = 0.71.

Holotype. ♂ ( SEHU), “ Ulu Gombak , Selangor, Malaysia, 8.xii.2013, M.J. Toda ”.

Distribution. Peninsular Malaysia.

Remarks. This species resembles P. afoliolata in having the small postgonites ( Fig. 3 F–H), but can be distinguished from it by the diagnostic characters.

Etymology. Referring to the lack of free sclerite within aedeagal outer membrane.

Figures

FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H.

FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm.