Phortica (Ashima) afoliolata, Toda & Bänziger & Sati & Fartyal & Suwito & Katoh, 2020

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 10

publication ID

https://doi.org/ 10.11646/zootaxa.4789.1.1

publication LSID

lsid:zoobank.org:pub:311A27A2-A053-4D20-A43E-EF57DBE55166

DOI

https://doi.org/10.5281/zenodo.5221268

persistent identifier

https://treatment.plazi.org/id/03C86C16-EB2F-5753-FF6E-DFF4FEC3F811

treatment provided by

Plazi

scientific name

Phortica (Ashima) afoliolata
status

 

1. Phortica (Ashima) afoliolata View in CoL species group, new

Diagnosis. Aedeagal, median rod distally diffentiated into two strongly sclerotized, asymmetric pieces (ch.58-4; Fig. 3F View FIGURE 3 ); arista with 4 or more ventral branches (ch.6-0; Fig. 2A View FIGURE 2 ); hypandrial phragma broader than lateral gonocoxites (ch.41-0; Fig. 3F View FIGURE 3 ).

Shared characters. Some interfrontal setulae as long as lowest setulae on ocellar triangle (ch.3-0). Arista apically without leaf-like expansion (ch.7-0); longest, dorsal branch longer than longest seta on pedicel ( Fig. 2A View FIGURE 2 ). Cibarial, medial sensilla all trichoid, apically pointed (ch.11-0; Fig. 2C View FIGURE 2 ). Prementum without pubescent patches (ch.12-0). Pseudotracheae in labellum 6 per side (ch.14-2; Fig. 2D View FIGURE 2 ). Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Mid and hind femora with gray patch submedially or largely dark brown (ch.16-1; Fig. 2G View FIGURE 2 ); all tibia with dark gray ring(s) or largely dark brown (ch.17,18,19-1; Fig. 2G View FIGURE 2 ). Apical, downward curved setula on fore tarsomere V neither thicker than surrounding setulae nor longer than tarsomere V (ch.20-0; Fig. 2E View FIGURE 2 ). Caudal bands on tergites III and IV medially interrupted/constricted (ch.21-2; Fig. 2F View FIGURE 2 ). Abdominal sternites III–V longer than wide; III not wider than IV; setulae on medial surface of III–V not so dense (ch.22–24,26–28-0; Fig. 2H View FIGURE 2 ). Epandrium widely round on ventral margin (ch.31-1; Fig. 3D View FIGURE 3 ); setae on and/or near ventral margin of epandrium not denser than those on other portions (ch.32-0; Fig. 3D View FIGURE 3 ), longer than others (ch.33- 1; Fig. 3D View FIGURE 3 ). Surstylus without expansion at caudoventral corner (ch.35-0; Fig. 3E View FIGURE 3 ), with peg-like teeth (ch.37-0; Fig. 3E View FIGURE 3 ). Hypandrium symmetric (ch.40-0; Fig. 3F View FIGURE 3 ). Pregonite ( Fig. 3G,H View FIGURE 3 ) partly strongly sclerotized (ch.43-0), rod-like, with apical knob (ch.44-1), apically narrowing basal extension nearly rectangular to main body (ch.49-1) and sensilla on submedial and basal portions (ch.52,53-1) but no serration on ventrodistal portion (ch.50-0); apical knob without fine serrations (ch.45-0), ventrosubapically protruded, forming apically pointed barbule (ch.46,47-0). Aedeagal, outer membrane bearing neither spiny ornamentation nor apical, long, narrow tube (ch.55-1, ch.57-0; Fig. 3 View FIGURE 3 F–H). Posterior, rod-like process of phallapodeme not longer than aedeagal median rod (ch.62-0; Fig. 3G,H View FIGURE 3 ). Aedeagal sheath and/or postgonites asymmetric (ch.63-1; Fig. 3F View FIGURE 3 ). Aedeagal sheath without lateral lobe(s) bearing strongly sclerotized postgonites (ch.65-0; Fig. 3G,H View FIGURE 3 ).

Included species. Phortica (Ashima) afoliolata ; P. (A.) eseta Chen ; P. (A.) qingsongi An & Chen*; P. (A.) xishuangbanna Cheng & Chen ; and P. (A.) efragmentata sp. nov. * The assignment was judged from the original description by An et al. (2015).

Remarks. This species group first branched off in the subgenus Ashima ( Fig. 1 View FIGURE 1 ), retaining a number of plesiomorphic characters such as the plumose (with 5 and 4 or more dorsal and ventral branches, respectively; ch.5,6-0) arista but lacking the apical, leaf-like expansion (ch.7-0), cibarial, medial sensilla all trichoid, apically pointed (ch.11-0), the fore tarsomere V apically without specially developed, downward curved setula (ch.20-0), and the abdominal sternites III–V small, not so conspicuously differentiated in shape and setation from each other (ch.22– 24,26–28-0).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Drosophilidae

Genus

Phortica

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