Megaphyllum projectum deubeli (Verhoeff, 1899)

Lazányi, Eszter & Vagalinski, Boyan, 2013, Redefinition of the millipede subgenus Megaphyllum sensu stricto Verhoeff, 1894 and neotype designation for Megaphyllum austriacum (Latzel, 1884) (Myriapoda: Diplopoda: Julida: Julidae), Zootaxa 3741 (1), pp. 55-100 : 62

Megaphyllum projectum deubeli (Verhoeff, 1899)

Figs 4a–d

Brachyiulus (Chromatoiulus) projectus deubeli Verhoeff, 1899 : Verhoeff 1899a: 192–193, Figs 24–25. Brachyiulus projectus deubeli: Verhoeff 1907: 303 .

Megaphyllum projectum deubeli: Lazányi & Korsós 2011: 49 –50, Figs 9–13 , 17c .

Material examined. Romania: 1♂, gonopods, slide preparation, “Rareul”, leg. “Hrevhan”, det. Attems (NHMW); 1♂, gonopods, slide preparation, Rodnageb. [Rodna Mts], Kuhhorn 3 [Ineu] (NHMW).

Distribution. Romania: Rodna Mts, Ineu peak (type locality) (Verhoeff 1899a).

Diagnosis. The smallest and most deviating subspecies of M. projectum . Differs from the other two subspecies by the broad and enlarged posterior process of the opisthomere (pp) ( Figs 4a–b, 4d ) and the short, ridge-like anterior process (ap) ( Fig. 4d ).

Solenomere processes (asp and psp) more or less equal in length.

Body length and height: male: 23.8mm, (21–28mm according to Verhoeff 1907), 1.7mm; females 29.9– 30.1mm, 2.5–2.7mm.

Remarks. Rare, strict endemic, its status has been clarified by Lazányi & Korsós (2011).

Figures

FIGURES 4 a – l. Megaphyllum projectum. (4 a – d) M. p. deubeli male from Băile Borşa, Maramureş Mts, Romania: right gonopods, (4 a) posterior and (4 b) mesal views; (4 c) left promere, meso-posterior view; (4 d) left opisthomere, lateral view. (4 e and 4 g – i) Megaphyllum projectum kochi male from Bukova, Malé Karpaty, Slovakia: (4 e) gonopods, posterior view; penis, (4 g) posterior and (4 h) right lateral views. (4 i) M. p. kochi female from Sanssouci, Germany: right vulva, posterior view (setae omitted). (4 f and 4 j – l) M. p. projectum male [originally Julus austriacus syntype, ZMB 1846]: (4 f) right gonopods, lateral view; (4 j) gonopods, posterior view; (4 k) right gonopods, mesal view. (4 l) M. p. projectum female from Haragistya, Aggtelek National Park, Hungary: right vulva, posterior view (setae omitted). Abbreviations: ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, sl: serrated lobe, P: promere. Scale bars: 0.2 mm.

FIGURES 9 a – g. Megaphyllum dentatum males and a female from Ljuboten, Serbia, Inv. No. 8018: (9 a) gonopods, posterior view; (9 b) right gonopods, mesal view; (9 c) right promere, meso-posterior view; (9 d) right opisthomere, lateral view; penis, (9 e) posterior and (9 f) left lateral views; (9 g) right vulva, anterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, ph: posterior hump, pp: posterior process, pr: protrusion, psp: posterior process of the solenomere, sl: serrated lobe, P: promere. Scale bars: 0.2 mm.

FIGURES 10 a – g. Megaphyllum glossulifer males and a female from Rhodopi Mts, Bulgaria: (10 a) gonopods, posterior view; (10 b) left gonopods, mesal view; (10 c) left promere, meso-posterior view; (10 d) left opisthomere, lateral view; penis, (10 e) posterior and (10 f) left lateral views; (10 g) left vulva, meso-posterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm.

FIGURES 11 a – g. Megaphyllum monticola. Males from Korakas Mts, Greece, ZMB 1300: (11 a) gonopods, posterior view and (11 b) right gonopods, mesal view and (11 c) left promere, meso-posterior view. Males from Ostrovo, Greece, Inv. No. 8019: (11 d) Right opisthomere, lateral view, penis, (11 e) posterior and (11 f) right lateral views, (11 f *) another, bent penis, left lateral view. (11 g) Left vulva, posterior view, female from Korakas Mts, Greece (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; op: operculum, pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm.

FIGURES 12 a – g. Megaphyllum bosniense. Syntype male from Bosnia, ZMB 2625: (12 a) gonopods, posterior view; (12 b) left gonopods, mesal view; (12 c) right promere, meso-posterior view; (12 d) left opisthomere, meso-posterior view. Male from Treskavac Mts, Montenegro: penis, (12 e) posterior and (12 f) right lateral views. Female from Maljen Mts, Serbia: (12 g) left vulva, meso-posterior view (setae omitted). Abbreviations: asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; pa: posterior ampulla, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm.

FIGURES 13 a – f. Megaphyllum rosenauense. Syntype male from Transylvania, Romania, ZMB 2622: (13 a) gonopods, posterior view; (13 b) right gonopods, mesal view; (13 c) right promere, posterior view; (13 d) right opisthomere, lateral view. Male from Miercurea Ciuc, Romania: penis, (13 e) posterior and (13 f) right lateral views. Abbreviations: asp: anterior process of the solenomere, ph: posterior hump, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm.

FIGURES 17 a – g. Megaphyllum sjaelandicum male and female from Dorpat, Estonia, ZMB 8734: (17 a) gonopods, posterior view; (17 b) left gonopods, mesal view; (17 c) right promere, meso-posterior view; (17 d) right opisthomere, mesal view; penis, (17 e) posterior and right lateral views; (17 g) right vulva, antero-mesal view (setae omitted). Abbreviations: ah: anterior hump; ap: anterior process, asp: anterior process of the solenomere, ca: central ampulla, ct: connecting tube; h: hump, l: lobe, op: operculum, pa: posterior ampulla, pp: posterior process, psp: posterior process of the solenomere, P: promere. Scale bars: 0.2 mm.