Peromelissa thoracites (Haeckel) Matsuzaki et al., 2015

Trubovitz, Sarah, Renaudie, Johan, Lazarus, David & Noble, Paula, 2022, Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific, Zootaxa 5160 (1), pp. 1-158 : 77-78

Peromelissa thoracites (Haeckel) Matsuzaki et al., 2015

Plate 38, Figs. 7A – 10 .

Lithomelissa thoracites n. sp., Haeckel, 1862, pl. 6, figs. 2–8.

Lithomelissa thoracites Haeckel, Hertwig, 1879, pl. 8, fig. 1.

Psilomelissa tricuspidata n. sp., Popofsky, 1908, p. 284, pl. 32, fig. 9; pl. 33, fig. 8.

Lithomelissa thoracites Haeckel, Popofsky, 1913 , text-figs. 44–47.

Lithomelissa thoracites Haeckel, Schröder, 1914 , text-figs. 61–62.

Lithomelissa thoracites Haeckel, Benson, 1966 , pl. 24, figs 10–12 (non fig. 13).

Dimelissa thoracites Haeckel, Petrushevskaya, 1971 , pl. 69, figs. 7–8.

? Dimelissa cf. thoracites Haeckel, Petrushevskaya, 1971 , pl. 69, figs. 5–6.

Lithomelissa monoceras Popofsky, Casey, 1971 , pl. 23.2, fig. 16.

Lithomelissa monoceras Popofsky, Renz, 1974 , pl. 18, fig. 14.

Lithomelissa monoceras Popofsky, Renz, 1976 , pl. 8, fig. 12.

Peromelissa phalacra Haeckel, Takahashi and Honjo, 1981 , pl. 7, fig. 3 (non figs. 4–5).

Lithomelissa thoracites Haeckel, Molina-Cruz, 1982 , pl. 4, figs. 12–13.

Peromelissa phalacra Haeckel, Nishimura and Yamauchi, 1984 , pl. 32, fig. 1 (non fig. 12).

Lithomelissa thoracites Haeckel, Nishimura and Yamauchi, 1984 , pl. 32, figs. 2–3.

non Lithomelissa cf. thoracites Haeckel, Nishimura and Yamauchi, 1984 , pl. 32, fig. 4.

Lithomelissa thoracites Haeckel, Boltovskoy and Riedel, 1987 , pl. 4, fig. 7.

Peromelissa thoracites forma thoracites, Van de Paverd, 1995 , pl. 66, figs. 9–10.

Peromelissa thoracites forma scaphopdium, Van de Paverd, 1995, pl. 65, figs. 17–21.

non Peromelissa thoracites?, Van de Paverd, 1995 , pl. 65, figs. 12–14, 16.

non Lithomelissa thoracites Haeckel, Chen and Tan, 1996 , pl. 28, figs. 1–4

Lithomelissa monoceras Popofsky, Chen and Tan, 1996 , pl. 28, fig. 5 (?fig. 6).

Peromelissa phalacra Haeckel, Sashida and Kurihara, 1999 , fig. 7.12.

Lithomelissa monoceras Popofsky, Tan and Su, 2003 , text-fig. 105.

non Dimelissa thoracites Haeckel group, Okazaki et al., 2005, fig. 12.15.

non Dimelissa thoracites Haeckel, Itaki, 2009 , pl. 15, figs. 1–8.

Dimelissa aff. thoracites Haeckel, Itaki, 2009 , pl. 15, figs. 14–16.

? Lithomelissa thoracites Haeckel, Sono et al., 2009 , pl. 3, fig. 3.

Dimelissa thoracites Haeckel, Itaki et al., 2010 , fig. 6.9–6.10.

Dimelissa monoceras Popofsky, Matsuzaki et al., 2014 , pl. 3, fig. 5.

Peromelissa thoracites Haeckel, Matsuzaki et al., 2015 , fig. 7.16.

Peromelissa tricuspidata Popofsky, Matsuzaki et al., 2015 , fig. 7.17.

non Lithomelissa thoracites Haeckel, Chen et al., 2017 , figs. 9–10, 12–13 (?figs. 11, 14).

Lithomelissa thoracites Haeckel, Trubovitz et al., 2020 , supplementary data 7.

Lithomelissa cf. thoracites Haeckel, Trubovitz et al., 2020 , supplementary data 7.

Remarks. Lithomelissa thoracites (Haeckel) was designated the type species of Dimelissa ( Campbell, 1951) by Petrushevskaya (1971). However, Matsuzaki et al. (2015) considered Dimelissa as a junior synonym of Peromelissa ( Haeckel 1881) and thus transferred L. thoracites Haeckel to the genus Peromelissa . Van de Paverd (1995) also suggested that this species should be considered Peromelissa , but the transfer was not formalized. Here we agree with Matsuzaki et al. (2015) that Dimelissa and Peromelissa should not be separate genera, and adopt their transfer here. However, the reason this species does not fit the definition of Lophophaena is not clear. Peromelissa thoracites (Haeckel) Matsuzaki et al. has often been confused with Peromelissa phalacra Haeckel in the literature. Like Peromelissa phalacra , this species has also sometimes been called Lithomelissa monoceras Popofsky. However , Lithomelissa monoceras as originally described and illustrated by Popofsky(1913) is a junior synonym of Peromelissa phalacra Haeckel. Subsequent authors (i.e., Casey, 1971, Renz, 1974, 1976) have illustrated specimens under the name Lithomelissa monoceras Popofsky that have a strong apical spine embedded in the cephalis wall, relatively large pores on the cephalis, and a relatively broad thorax, which are more characteristic of Peromelissa thoracites Haeckel than Peromelissa phalacra Haeckel , and are thus included in our synonymy.

Range. Late Upper Miocene—Recent ( Table 1 ).

Figures

FIGURE 7. Internal skeletal structure of Arachnocorys umbellifera (Haeckel, 1862) Petrushevskaya, 1971.

FIGURE 10. Internal skeletal structure of Lithomelissa alkonost n. sp.