Transennatia gratiosa (Waagen, 1884)

Chen, Zhong-Qiang, Campi, Monica J., Shi, Guang R. & Kaiho, Kunio, 2005, Post-extinction brachiopod faunas from the Late Permian Wuchiapingian coal series of South China, Acta Palaeontologica Polonica 50 (2), pp. 343-363 : 354

Transennatia gratiosa (Waagen, 1884)

Figs. 10E–H , 11 .

Productus gratiosus n. sp.; Waagen 1884: 691, pl. 72: 3–7.

Productus gratiosus Waagen ; Diener 1897: 23, pl. 3: 3–7; Mansuy 1913: 115, pl. 13: 1a, b; Broili 1916: 12, pl. 116: 4, 5, 7–11; Chao 1927: 44, pl. 4: 6–8; Chi−Thuan 1962: 491, pl. 2: 5–7.

Productus (Dictyoclostus) gratiosus Waagen ; Huang 1932: 32–33, pl. 2: 3; 1933: 88, pl. 11: 14a, b.

Marginifera gratiosa (Waagen) ; Licharew 1936: 60, 118, pl. 4: 11–19; Reed 1944: 98, pl. 19: 6, 7.

Dictyoclostus gratiosus (Waagen) ; Zhang and Ching (Jin) 1961: 411, pl. 4: 12–18; Wang et al. 1964: 291, pl. 45: 14–19.

Transennatia gratiosa (Waagen) ; Waterhouse 1975: 10–11.

Gratiosina gratiosa (Waagen) ; Grant 1976: pl. 33: 19–26; Licharew and Kotlyar 1978: pl. 12: 5, 6; pl. 12: 1a, b.

Asioproductus gratiosus (Waagen) ; Yang et al. 1977: 350, pl. 140: 5a–c; Feng and Jiang 1978: 254, pl. 90: 1, 2; Tong 1978: 228, pl. 80: 7a, b; Li et al. 1980: 373, pl. 164: 14a–c; pl. 166: 5, 6.

Asioproductus bellus sp. nov.; Chan (Zhan) 1979: 85–86, pl. 7: 7–13, 9: 8–10.

Transennatia gratiosa (Waagen) ; Liao 1980b: pl. 3: 25; Liu et al. 1982: 185, pl. 132: 9a–d; Wang et al. 1982: 214, pl. 92: 6–8; pl. 102: 4–9; Yang 1984: 219, pl. 33: 7a–c; Zeng et al. 1995: pl. 5: 14, 15; Tazawa and Matsumoto 1998: 6, pl. 1: 4–8; Tazawa et al. 2000: 7, pl. 1: 3–5; Tazawa and Ibaraki 2001: 7, pl. 1: 1–3.

Locality and horizon: Daijiagou section; 4 th and 5 th members of the adductor scars

Lungtan Formation; Chuanmu section; 5 th member of the Lungtan Formation.

Material.—NMV P309636–639.

Remarks.—A restudy ( Grant 1976) of the topotype specimens of Productus gratiosus Waagen, 1884 from the Wargal and Chhidru Formations at Chhidru and Jabbi of the Salt Range, Pakistan allows us to understand the internal features as following: the dorsal valve has a sessile W−shaped cardinal process; a pair of elongate, highly raised adductor scars are separated by a low, thin median septum; the ventral valve is tuberculate, and embraces an elongate, highly raised median adductor scar; the diductor scars are wide, flabellate and deeply grooved on the floor of each side of the scar platform. These features are confirmed in our specimen ( Fig. 11 ). This Salt Range species has also been referred to as the type species of Asioproductus Zhan (in Yang et al. 1977). Later, Zhan (1979) altered his specimens to Asioproductus bellus Zhan, 1979 , but did not compare his new material with Waagen’s species. Liang (1990: 162) differentiated Asioproductus from Transennatia by the appearance of a Marginifera −type, bilobate cardinal process. However, one of Zhan’s (1979: pl. 6: 9) illustrations clearly shows a W−shaped cardinal process. As a consequence, Asioproductus is synonymous to Transennatia . Another species Transennatia insulpta Grant, 1976 (see also Chen 2004) can be distinguished from the type species by its evidently smaller size and more extended ears.

Figures

Fig. 10. A. Meekella kueichowensis (Huang, 1933), dorsal (A1), ventral (A2), and lateral (A3) views of NMV P309622. B. Chengxianoproductus cf. changxingensis Liao and Meng, 1986, ventral (B1), posterior (B2), lateral (B3), and dorsal (B4) views of NMV P309640 (B); B5, enlargement of fine, densely arranged spine bases of B1. C, D. Spinomarginifera alpha Huang, 1932, lateral (C1), posterior (C2), ventral (C3), and dorsal (D) views of a complete specimen NMV P309643 and a dorsal external mold NMV P309644. E–H. Transennatia gratiosa (Waagen, 1884), ventral (E1, G, H1), anteroventral (E2, H2), posterior (E3, H3), lateral (E4, H4), and internal (F) views of a ventral internal mold NMV P309639 (E), and latex replica NMV P309638 (F) of the ventral internal mold (E –E), a ventral internal mold NMV P309636 (G) and a dorsal external mold NMV P309637 (H –H). A, E, F, G, from the 4th member of the Lungtan For1 4 1 4 mation of Daijiagou; B, from the 1st member of the Lungtan Formation of Daijiagou; C, D, H, from the 5th member of the Lungtan Formation of Daijiagou.

Fig. 11. Ventral interior of Transennatia gratiosa (Waagen, 1884); based on a ventral internal mold NMV P309639.