Chilicola (Oroediscelis) goloboffi Packer and Dumesh, 2019

Packer, Laurence & Dumesh, Sheila, 2019, Fifteen new species of Chilicola (Oroediscelis) (Hymenoptera: Colletidae: Xeromelissinae) with illustrated keys to the males and females of the subgenus, Zootaxa 4559 (1), pp. 1-56 : 11-13

Chilicola (Oroediscelis) goloboffi Packer and Dumesh , new species

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( Figs. 24–30 , 79 , 85 , 147 )

Diagnosis. The male of this species can be differentiated from all other species of Chilicola by the combination of: S5 with conical process on each side and S4 with elongate process not strongly upcurved. The only other species with a process on S5, C. abrebotellas , has the processes on S4 strongly curved. The female can be differentiated by the following combination: malar space greater than 3/4 width (at most 4/5 as long as wide); head 1.2X as long as wide and large body size (7.5 mm in length). The most similar species are C. carpenteri and C. rozeni , but both are much smaller (5.6–5.8 mm in length) and have a longer head (>1.25X as long as wide).

Description. Male: Length 8.0 mm; forewing length 5.5 mm, head width 1.7 mm.

Head: about 1.1 X as long as wide, 94:85. Clypeus with transverse apical yellow band, paraocular area yellow to just above level of dorsal margin of clypeus, more elongate medially, upper margin acute; mandible with basal yellow maculation. Clypeus and supraclypeal area imbricate, punctures small, i=1–4 d on clypeus, i=0.5–3 d on supraclypeal area; lower paraocular area with large irregular punctures, i=0.2–4 d; frontal area punctures large, crowded. Malar space approximately half as long as wide (7:13); lower ocular tangent approximately at mid length of clypeus. Longest hairs of face above antennal sockets,>2.3 MOD, and on vertex, 2.0–2.2 MOD; genal beard with hairs up to 4 MOD. F1 as long as F2 on anterior surface (13:13) but longer on posterior surface (16:13); F2 with subapical seta subequal in length to F3, F3 with seta <0.5X length of F4. Mesosoma: pronotum with dense punctures, i<d; mesoscutum and mesoscutellum imbricate, dull, punctures distinct, i <d; metanotum less regularly punctate, i<3d; mesepisternum punctures large and dense, i = 0.5–1.5 d. Metepisternum strongly rugulose above, weakly wrinkled below, posteriorly shiny; metapostnotum shorter than mesoscutellum (20:22) with irregular, anastomosing, sometimes incomplete, longitudinal striae; lateral surface of propodeum imbricate, punctures large, shallow, longitudinally effaced, becoming rugose posteroventrally. Longest hairs of thorax on mesepisternum ~3 MOD, most hairs of mesoscutum ~1.5 MOD. Stigma much shorter than marginal cell on wing margin (35:55); marginal cell abruptly set off from leading edge of forewing, apex almost truncate; distal stigmal perpendicular slightly basal to first submarginal crossvein. Mesofemur weakly concave basoventrally. Metafemur with posteroventral margin sharp and weakly convex for apical 2/3; metatibia 3.3 X as long as greatest depth (66:20) which is just basal to dorsally incomplete subapical brush; bare area of ventral surface shining, without microsculpture; posterior crest sinuate, ending in a strong, acute tooth; anterior crest transverse, short, evenly rounded in outer view. Metabasitarsus less than 3/4 as long as remaining tarsomeres combined (43:60), ventral process strong, positioned at basal 1/4 of basitarsal length; anterior process swollen basally, apex forming acute tooth.

Metasoma: dull due to dense microsculpture except on submarginal areas which are shiny; T1 punctures dense on disc, i <d, sparser elsewhere, apex of apical impressed area impunctate for ~0.5 MOD; punctures increasingly sparse on succeeding apical impressed areas; apicolateral patches of white, plumose hairs on T1–T3. S1 erect hairs longest posterolaterally, 2 MOD; S2 erect hairs longest anteriorly, <2 MOD; S3 with few long hairs posterolaterally, <2 MOD; S4 and S5 erect hairs sinuate, 3 MOD; S6 apical tuft ~2.5 MOD. S1 and S2 unmodified; S3 with longitudinal sublateral boss; S4 process long, 3 MOD, broad, ventral margin sinuate, apex concave in lateral view, space between processes narrow, parallel-sided; S5 process short; S6 apicolateral fovea almost 3X as long as greatest width, disc concave, apical impressed area distinct, elongate ~1.1 MOD, subapical setal row long, ~1.5 MOD, at ~40° to impressed area. S5 and S6 oriented at angle of ~45° to S1–S4.

Terminalia : refer to Figs. 26–28 .

Female: As in male except for usual secondary sexual characteristics and as follows: Length 7.5 mm, forewing length 4.6 mm, head width 1.7 mm. Face and mandibles black. Punctures finer and more widely spaced on frontal area; facial fovea shiny, punctures sparse, minute; mesepisternum punctation finer and sparser, i=1.5–2 d; metasomal terga with punctures shallow, small, and much sparser, i>3d; apical impressed areas with minute punctures on T1–T2 laterally, impunctate on T3–T5. Genal area shiny; mesoscutum anteriorly shiny, not as strongly imbricate; metapostnotum with striae stronger and more developed; metasomal terga shinier.

Material studied. Holotype male: ARGENTINA, Jujuy, Tumbaya , 24.i.1948, Monrós & Willink [ LACM] . Paratype female: ARGENTINA, Salta, El Alisal, Fritz, iv.1991, [ AMNH]. The holotype bears a LACM GUI— ENT 240111 . The holotype is in good condition except for some “dirt” on the dorsal surface of the mesosoma and the apex of the right forewing missing. The paratype is in good condition except for the left wing, which is torn basally. [See Figs. 184–185 for distribution map] .

Etymology. This species is named in honour of Dr. Pablo Goloboff of the Instituto Miguel Lillo in Tucumán, a well-known researcher into, and teacher of, phylogenetic methodology and a computer programmer of phylogenetic software par excellence, in gratitude for his assistance with the first author’s fieldwork in Argentina in February of 2003.

Comments. The two apical external sterna are upturned in the male, though not as strongly as in the preceding species.

Figures

FIGURES 24–30. Chilicola goloboffi. 24. male lateral habitus (holotype); 25. male face (holotype); 26. male genital capsule, dorsal view (holotype); 27. male S7, dorsal view (holotype); 28. male S8, dorsal view (holotype); 29. female lateral habitus (paratype); 30. female face (paratype).

FIGURES 79–80. Ventral view of S4 of male to show length (red lines) of processes in relation to distance separating them (blue lines). 79. length> distance between processes (C. goloboffi holotype); 80. length at most 2/3 distance separating processes (C. maculiPes paratype).

FIGURES 85–86. Lateral view of metasomal apex to show: 85. S4 process straight and apex entire (C. goloboffi holotype); 86. notched apex to process of S4 (C. fritzi holotype).

FIGURES 147–148. Head, frontal view, to show relative length to width. 147. L~1.2XW (C. goloboffi paratype); 148. L>1.25W (C. carPenteri paratype).

FIGURES 184–185. Distribution maps of newly described Chilicola (Oroediscelis) species. 184. Large scale map of South America showing distribution of new species; 185. Small scale map of northern Argentina to show distribution within the country.