Sinoennea Kobelt, 1904

Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Sajan, Sheikh, Tripathy, Basudev & Chen, Zhe-Yu, 2020, A review of the Diapheridae (Gastropoda: Eupulmonata: Streptaxoidea), with special emphasis on India and Myanmar, Raffles Bulletin of Zoology 68, pp. 682-718 : 704

Sinoennea Kobelt, 1904

Ennea (Sinoennea) Kobelt, 1904a: 27 .

Indoennea Kobelt, 1904a: 28 .

Sinoennea — Peile, 1935: 382. (synonymised Indoennea with Sinoennea )

Sinoennea — Zilch, 1960: 573. (including the subgenera Indoennea and Sinoennea )

Sinoennea — Richardson, 1988: 154.

Indoennea — Schileyko, 2000: 799.

Sinoennea — Schileyko, 2000: 800.

Type species. Ennea strophiodes Gredler, 1881 ( Fig. 14A–E ), by original designation.

Diagnosis. Shell usually elongate ovoid and cylindrical, glossy, and regularly ribbed. Apex usually domed. Basal swelling behind peristome usually present. Parietal lamella long, well-developed, divided to internal and external portions that usually differ in orientation. Palatal wall with an upper palatal tooth that sits on the peristome edge and faces the parietal lamella, and a lower palatal tooth that is situated deeper and corresponds with a depression behind the peristome. Small basal tooth sometimes present. Columellar lamella usually deeply situated, sometimes corresponding with a depression on the umbilical side ( Fig. 1 ). Umbilicus open or closed, periumbilical keel might be present.

Differential diagnosis. Bruggennea agrees with Sinoennea in terms of apertural dentition, but possesses varices (remaining expanded peristomes of juvenile shells) that are imperfectly arranged in two rows on opposite sides of the spire. Parasinoennea , new genus, differs from Sinoennea mostly in the triangular aperture shape and the adnate parietal callus; other differences listed under the genus. Platylennea , new genus, differs from Sinoennea in the general shell shape (aciculid-like, elongate conical), the weak parietal callus, and the differently arranged apertural barriers (see under the genus). Rowsonia , new genus, differs from Sinoennea in the slender conical shell shape, the presence of deeply situated parietal/basal folds, and the basal swelling and the depression corresponding with the lower palatal tooth are absent. Pupennea , new genus, mostly differs from Sinoennea species by the short parietal lamella, the pointed, small columellar lamella, the well-developed columellar denticle, and the weak parietal callus.

Distribution. From northern India to northern Borneo in the south and Japan in the east ( Fig. 10 ).

Remarks. This genus consists of over 80 species with diverse shell characters (shell size, sculpture, aperture formation). Still, at present we see no possibility to subdivide it in a useful way based on shell characters.

Indoennea was described by Kobelt (1904a, published on 22 February), and was later ( Kobelt, 1904b, published in May) again declared new. The type species ( Ennea blanfordiana Godwin-Austen, 1872 , Fig. 14F–J ), was designated to be the type species in the latter publication. Thus, the type species of Indoennea is fixed by subsequent designation.

Figures

Fig. 14. Shells of Sinoennea species. A–E, Ennea strophiodes Gredler, 1881 (type species of Sinoennea Kobelt, 1904), SMF 42769 (syntype); Sinoennea blanfordiana Godwin-Austen, 1872 (type species of Indoennea Kobelt, 1904), NHMUK 1903.7.1.2872 (syntype). Scale represents 5 mm.

Fig. 1. Nomenclature of the apertural barriers. A–H, Sinoennea; I, J, Pupennea, new genus; K, L, Sinoennea fartoidea; M, N, Platylennea, new genus; O, Rowsonia, new genus; P, Gulella (Huttonella). Abbreviations: BK, basal keel; BS, basal swelling; BT, basal tooth; CD, columellar denticle; CL, columellar lamella; CTd, depression corresponding with the columellar lamella; IBD, inner basal denticle; ICE, inner columellar edge; LBD, lower basal denticle; LPF, lower palatal fold; LPT, lower palatal tooth; LPTd, depression corresponding with the lower palatal tooth; MPF, middle palatal fold; OBD, outer basal denticle; PL, parietal lamella; PT, palatal tooth; S, sinulus; UBD, upper basal denticle; UPF, upper palatal fold; UPT, upper palatal tooth.

Fig. 10. Distribution of Parasinoennea, new genus, Sinoennea Kobelt, 1904, Pupennea, new genus, Platylennea, new genus, and Rowsonia, new genus.