Pupennea planguncula ( Benson, 1863 )

Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Sajan, Sheikh, Tripathy, Basudev & Chen, Zhe-Yu, 2020, A review of the Diapheridae (Gastropoda: Eupulmonata: Streptaxoidea), with special emphasis on India and Myanmar, Raffles Bulletin of Zoology 68, pp. 682-718 : 700

Pupennea planguncula ( Benson, 1863)

( Figs. 13 , 22D )

Pupa planguncula Benson, 1863: 426 . Ennea (Huttonella) planguncula — Nevill, 1878: 7. Ennea stenostoma W.T. Blanford, 1881: 208 , pl. 2, fig. 17. Ennea stenostoma var. minor W.T. Blanford, 1881: 208 , pl. 2, fig. 16. Ennea ( Indoennea ?) stenostoma — Kobelt, 1904b: 157, pl. 20,

fig. 19. Ennea (Huttonella) planguncula — W.T. Blanford & Godwin-

Austen, 1908: 23. (considered Ennea stenostoma as synonym) Ennea planguncula — Richardson, 1988: 25. Gulella (Huttonella) planguncula — Ramakrishna et al., 2010: 195. Sinoennea planguncula — Raheem et al., 2014: 145, fig. 95A, B.

Type locality. “in regione Orissae et prope fluvium Nerbuddae” ( S. planguncula ); “In montibus ‘Galconda’ dictis, haut procul ad urbe Vizagapatam” ( S. stenostoma ); “In montibus haut procul ad urbe Karnul (Karnool) Indiae meridionalis” ( S. stenostoma var. minor ).

Material examined. Lectotype (designated by Raheem et al. 2014, UMZC I.103310.B); 1 shell ( NHMUK 1906.1.1.292), no locality, coll. W.T. Blanford, figured in FBI [ Fauna of British India].

Diagnosis. Peristome discontinuous, parietal callus only indicated with a very slight calcareous layer. Basal swelling only slight. The depression on the outer side of the palatal wall is deep and corresponds with the large parietal tooth (lower parietal tooth). Parietal lamella elevated, well-developed, but short, with no inner and outer parts distinguishable. Palatal wall with a strongly developed bifurcated structure consisting of two parallel ridges entering the aperture from the peristome edge, and an incision between them which faces the tip of the parietal lamella. The upper denticle of this structure is probably homologous with the upper palatal tooth, whereas the lower is the lower palatal tooth. There is an additional, relatively slender, short, blunt tooth between the basal area and the complicated palatal tooth (outer basal denticle on Fig. 1 ). Another blunt basal tooth (inner basal tooth on Fig. 1 ) is situated in some distance from the lip, and relatively close to the columellar denticle. Columellar denticle large but blunt, situated on the columellar lip. Columellar lamella pointed, small, situated behind and little higher in position than the columellar denticle (in the parieto-columellar corner).

Figures

Fig. 13. Pupennea planguncula (Benson, 1863), NHMUK 1875.3.6.39. Both scales represent 1 mm; scale “a” refers to fig. A, scale “b” refers to figs. B–I.

Fig. 22. Apertures of Streptaxidae (A) and Diapheridae (B–P). A, Gulella (Huttonella) bicolor (Hutton, 1834); B, Rowsonia beddomei (W.T. Blanford, 1881); C, Platylennea subcostulata (W.T. Blanford, 1881); D, Pupennea planguncula (Benson, 1863); E, Diaphera polita, Páll-Gergely new species; F, Diaphera cylindrelloidea (Stoliczka, 1871); G, Bruggennea cf. bongi (Dance, 1970), H, Parasinoennea ovulum (Bavay & Dautzenberg, 1912); I, Sinoennea otostoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely, Reischütz, Maassen, Grego & Hunyadi, 2020; J, Sinoennea infantilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; K, Sinoennea variabilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; L, Sinoennea sutchariti Páll-Gergely & Hunyadi in Páll-Gergely et al., 2020; M, Sinoennea montawana Páll-Gergely & Hunyadi, new species; N, Sinoennea angustistoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely et al., 2020, O, Sinoennea woodthorpei (Peile, 1929); P, Sinoennea fartoidea (Theobald, 1870). Not to scale.

Fig. 1. Nomenclature of the apertural barriers. A–H, Sinoennea; I, J, Pupennea, new genus; K, L, Sinoennea fartoidea; M, N, Platylennea, new genus; O, Rowsonia, new genus; P, Gulella (Huttonella). Abbreviations: BK, basal keel; BS, basal swelling; BT, basal tooth; CD, columellar denticle; CL, columellar lamella; CTd, depression corresponding with the columellar lamella; IBD, inner basal denticle; ICE, inner columellar edge; LBD, lower basal denticle; LPF, lower palatal fold; LPT, lower palatal tooth; LPTd, depression corresponding with the lower palatal tooth; MPF, middle palatal fold; OBD, outer basal denticle; PL, parietal lamella; PT, palatal tooth; S, sinulus; UBD, upper basal denticle; UPF, upper palatal fold; UPT, upper palatal tooth.