Fetilinia dentator, Lowe & Kovařík, 2020

Lowe, Graeme & Kovařík, František, 2020, Fetilinia dentator gen. et sp. n. from Pakistan (Scorpiones: Buthidae), Euscorpius 328, pp. 1-10 : 4-10

Fetilinia dentator sp. n.

( Figs. 1–31 , Table 1 )

http: //zoobank. org/urn: lsid: zoobank. org: act: 2883418A- AB1A-4991-AE48-FFBB13854627

TYPE LOCALITY AND TYPE DEPOSITORY. Pakistan, Khyber Pakhtunkhwa (formerly North-Western Frontier) Province, Karak , 33.102°N 71.049°E ; FKCP.

TYPE MATERIAL EXAMINED. Pakistan, Khyber Pakhtunkhwa (formerly North-Western Frontier) Province, Karak , 33.102°N 71.049°E, 23 June 2010, 1♂ subadult (holotype) 1♀ juvenile (paratype), leg. Z. Ahmed , FKCP.

ETYMOLOGY. The specific epithet refers to the prominently enlarged dentition on the metasoma, and is chosen to rhyme with the specific epithet palpator of Kraepelinia .

DIAGNOSIS. See generic diagnosis.

DESCRIPTION (subadult ♂). The subadult male is 22 mm long. The habitus is shown in Figs. 1–2 and 30–31 . For position and distribution of trichobothria of pedipalps see Figs. 15–18 , and 20–21.

Coloration ( Figs. 1–2 , 6–9 ). The base color is uniformly yellow, with dark carinae indicated on the pedipalp femur and patella, and on metasoma III–V.

Carapace ( Figs. 10 , 25 ). Trapezoidal, anterior margin almost straight, with 10 marginal macrosetae; surface with dense, coarse granulation, weaker anteriorly; carinae absent; area between anterior median carinae smooth; median ocular tubercle smooth, eXcept for a few posterior granules; median eyes large, well separated; 5 lateral eyes (3 larger, 2 smaller). Chelicera ( Figs. 24–25 ). Fingers comply with the basic pattern of buthid dentition (Vachon, 1963); fiXed finger with large distal denticle, one subdistal denticle and two basal denticles fused into bicusp, two denticles on ventral surface; dorsal margin of movable finger with 5 denticles: one large distal denticle, medium-sized subdistal denticle, large medial denticle, and two small, partially fused basal denticles; ventral margin with 3 denticles: one large distal denticle, and two smaller denticles in medial and basal positions.

Mesosoma ( Figs. 10–12 ). Tergites I–VI tricarinate with a median carina, and a weak pair of lateral carinae; carinae coarsely granular, the lateral pair on tergites I–II rather indistinct; all carinae short with only ca. 2–5 granules, confined to posterior half or third of tergite; entire tergites covered with dense, coarse granulation; pretergites smooth; tergite VII pentacarinate, median carina a granulated hump, lateral carinae well developed, coarsely granular; sternites with smooth surfaces locally finely granulated, more so on sternite III and VII; sternite III–VI without carinae, VII with 2 pairs of weak to moderate, smooth to granulated carinae; sternal chaetotaxy: sternite III–VI posterior margins bearing 3 macrosetae, III bearing 5–7 macrosetae on medial surface, IV–VII with 2–4 macrosetae; pectines with margins eXtending to half of sternite IV in males, with 3 marginal lamellae, 6–8 middle lamellae; lamellae and fulcra bear numerous short, fine, dark macrosetae; pectine basal piece and genital opercula smooth with fine macrosetae; pectinal tooth count 23– 23 in male.

Metasoma and telson ( Figs. 3–9 ). Metasoma I–III with 10 granulated carinae, median lateral carinae on segments I–II complete, median lateral carinae on segment III incomplete, indicated by ca. 10–18 granules on posterior part; ventromedian and ventrolateral carinae stronger on segments II–III, with conspicuously enlarged, dentate granules increasing in size posteriorly; metasoma IV with 4, and metasoma V with 2–4 granulated or crenulated carinae, dorsolateral carinae granulated onIVandincompleteonV, ventrolateralcarinaestrong, irregularly crenulated on V with several enlarged, lobate granules that become larger posteriorly; ventromedian carinae of metasoma IV–V absent; intercarinal surfaces partly smooth and partly irregularly granulated, with granules mostly on dorsomedial and ventral surfaces; lateral anal arch divided into 3 lobes; ventral anal arch armed with a regular series of ca. 10 coarse granules; telson with distinctly elongated, smooth vesicle; aculeus robust, equal to or shorter than vesicle in length, moderately curved; subaculear tubercle absent; chaetotaxy: metasomal segments and telson sparsely setose; long macrosetae dispersed irregularly on lateral and ventral surfaces.

Pedipalps ( Figs. 13–23 ). Segments slender, with chelae narrower than patella; femur with 3 strong, granulated carinae: dorsoeXternal, dorsointernal, and ventrointernal, other carinae obsolete; dorsal, lateral and ventral surfaces smooth eXcept for a few small solitary granules, internal surface smooth eXcept for several coarse solitary granules; patella with 7 smooth, obsolete carinae; setation very sparse, with a few large solitary macrosetae; chela smooth, carinae obsolete, a few large macrosetae present; dentate margins of movable finger with 8 rows of granules, of fiXed finger with 7 rows of granules, each flanked by a single eXternal and internal accessory granule, distal ends of fingers with 5 subterminal granules; trichobothrial pattern type A-β with reductive neobothriotaXy (femur petite d 2 absent).

Legs ( Figs. 26–29 ). Legs with robust patellae, tibiae and tarsi; femora with a few solitary macrosetae; tibiae I–III with 3–6 long macrosetae in dorsal (retrosuperior) series, not forming a tibial ‘bristle comb’; basitarsi I–III compressed, with two irregular series of shorter ventral (proinferior and retroinferior) macrosetae, and a single linear series of longer dorsal macrosetae (forming a basitarsal ‘bristle comb’with 4–5 setae on leg I, 5 setae on leg II, 6 setae on leg III); leg IV without basitarsal compression, longer than legs I–III, leg I–IV femora and patella with indications of 4–6 carinae, which are usually obsolete; paired ventral carinae on femora granulate or denticulate; tibial spur on legs III–IV moderate; prolateral pedal spurs basally bifurcate; retrolateral pedal spurs simple; telotarsi with 2 ventral rows of fine macrosetae; ungues elongate, curved.

Measurements. See Table 1 .

Figures

Figures 1–2. Fetilinia dentator gen. et sp. n., subadult male holotype in dorsal (1) and ventral (2) views. Scale bar: 10 mm.

Figures 3–9. Fetilinia dentator gen. et sp. n., subadult male holotype, metasoma IV–V and telson ventral view (3), metasoma and telson dorsal (4, 7), ventral (5, 8), and lateral (6) views, metasoma V and telson lateral view (9). Figures 3–5 are under UV light. Scale bar: 10 mm (4–8).

Figures 10–23: Fetilinia dentator gen. et sp. n., subadult male holotype. Figures 10–12. Carapace and tergites (10) and sternopectinal area and sternites (11–12) Figures 13–23. Pedipalp segments, chela dorsal (14), eXternal (15), ventral (16) views, patella dorsal (17), dorsoeXternal (18), and ventral (19) views, femur and trochanter dorsoeXternal (20), internal (21), and ventral (22) views, movable (13) and fiXed (23) finger dentition. Trichobothrial pattern is indicated by white circles in Figures 15–18 and 20–21. Figures 10–11 under UV light.

Figures 24–29: Fetilinia dentator gen. et sp. n., subadult male holotype. Figures 24–25. Chelicerae ventral (24) and chelicerae dorsal with anterior part of carapace with median and lateral eyes (25). Figures 26–29. Left legs I–IV, retrolateral aspect (respectively).

Figures 30–31. Fetilinia dentator gen. et sp. n., subadult male holotype in dorsal (30) and ventral (31) views, under UV light.