Hoplopholcus atik, Huber, 2020

Huber, Bernhard A., 2020, Revision of the spider genus Hoplopholcus Kulczyński (Araneae, Pholcidae), Zootaxa 4726 (1), pp. 1-94: 72-75

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Hoplopholcus atik

sp. n.

Hoplopholcus atik   sp. n.

Figs 356–371 View FIGURES 356–361 View FIGURES 362–371

Pholcus opilionoides   (misidentification)— Roewer 1959: 10 (only specimens from “Atik, Amanus”).

Diagnosis. Distinguished from known congeners by shapes of procursus and bulbal processes ( Figs 362–363, 366–369 View FIGURES 362–371 ): tip of procursus with large dorsal process (arrow in Fig. 363 View FIGURES 362–371 ) and prolateral whitish protrusion (arrow in Fig. 362 View FIGURES 362–371 ), without sclerotized prolateral process or ridge, ventral spine of procursus proximally strongly curved, ventral ‘knee’ of procursus small but distinct; ventral bulbal sclerite distally strongly widened (in retrolateral view, Fig. 369 View FIGURES 362–371 ). Females are difficult to distinguish externally from congeners; dorsal arc with distinctive median ridge (arrow in Fig. 371 View FIGURES 362–371 ), median pouch of uterus externus poorly visible in uncleared specimens ( Fig. 359 View FIGURES 356–361 ); pair of lateral pouches of uterus externus connected to ventral arc, with posteriorly diverging sclerotized external margins ( Fig. 370 View FIGURES 362–371 ).

Type material. TURKEY, Hatay: ♂ holotype, NHMG, Amanus [ Amanus Mountains , Nur Dağlari], Atik [~ 36.52°N, 36.24°E], Roewer collection number “TQ 139” (“ Aatik, Grotte I ”), 17.vi.1953 (K. Lindberg) GoogleMaps   .

Other material examined. TURKEY: 3♂ 8♀, NHMG and   1♂ 2♀, ZFMK (Ar 20966), same data as holotype GoogleMaps   .

Uncertain identity (see Description below). TURKEY, Erzurum: 1♂ 1♀ 1 juv., NMNHS, Tercan District, small cave near road at 2 km W Tercan (39.773°N, 40.356°E), 1465 m a.s.l., under stones and on walls, 18.vii.2006 (P. Stoev, S. Lazarov).

Etymology. The species name is derived from the type locality; noun in apposition.

Description. Male (holotype). MEASUREMENTS. Total length 4.7, carapace width 1.7. Distance PME-PME 130 µm; diameter PME 110 µm; distance PME-ALE 40 µm; diameter AME 60 µm; distance AME-AME 30 µm. Leg 1: 37.4 (10.2 + 0.8 + 10.2 + 13.5 + 2.7), tibia 2: 6.7, tibia 3: 5.2, tibia 4: 6.2; tibia 1 L/d: 57.

COLOR (in ethanol). Carapace mostly ochre-yellow, ocular area and clypeus light brown; sternum ochre-yellow, medially slightly darker and margins light brown; legs ochre-yellow, without darker rings; abdomen monochromous gray, only dorso-posteriorly with indistinct darker marks; ventrally with light brown plate in front of gonopore.

BODY. Habitus similar to H. labyrinthi   (cf. Figs 6–7 View FIGURES 6–11 ). Ocular area slightly elevated. Deep thoracic pit and indistinct pair of shallow furrows diverging from pit toward posterior margin. Clypeus unmodified. Sternum wider than long (1.3/0.8), unmodified. Abdomen oval, dorso-posteriorly rounded.

CHELICERAE. As in Figs 364–365 View FIGURES 362–371 , with pair of latero-distal apophyses provided with two modified coneshaped hairs each; fine stridulatory ridges barely visible in dissecting microscope.

PALPS. As in Figs 356–358 View FIGURES 356–361 ; coxa with retrolateral bulge, trochanter barely protruding ventrally, femur with retrolateral dark line and prolateral stridulatory pick; procursus ( Figs 362–363 View FIGURES 362–371 ) with small but distinct ventral ‘knee’, with distinctive dorsal process and prolateral whitish protrusion (arrows in Figs 362, 363 View FIGURES 362–371 ), ventral spine proximally strongly curved, without prolateral process or ridge; genital bulb ( Figs 366–369 View FIGURES 362–371 ) with distally strongly widened (in retrolateral view) ventral sclerite; dorsal membranous process distinct ( Fig. 367 View FIGURES 362–371 ).

LEGS. Femur 1 with single row of ~25 ventral spines; with curved hairs (but most hairs missing); few vertical hairs; retrolateral trichobothrium of tibia 1 at 5%; prolateral trichobothrium present on all leg tibiae; tarsi without distinct pseudosegments but with many small platelets.

Male (variation). Tibia 1 length in three other males: 10.2, 11.2, 11.5. Most males with two modified hairs on each cheliceral apophysis, one male with only one hair on each side. Largest males with some stronger hairs also on femur 2 (ventro-distally). Single male from Erzurum Province very similar to holotype but dorsal process on procursus smaller; tibia 1: 9.1.

Female. In general similar to male but without spines on legs. Tibia 1 in five females (except female from Erzurum Province): 10.0–11.4 (mean 10.5).

FEMALE GENITALIA. Epigynum as in Fig. 359 View FIGURES 356–361 , triangular to bell-shaped; internal median pouch of uterus externus, sclerotized arcs, and lateral pouches usually visible in uncleared specimens; with pair of light brown weakly bulging areas in front of epigynum and simple, very short but wide posterior plate. Internal genitalia with distinctive median ridge at dorsal sclerotized arc (arrow in Fig. 371 View FIGURES 362–371 ), low and indistinct median pouch of uterus externus, pair of lateral pouches of uterus externus connected anteriorly to ventral arc and posteriorly with diverging sclerotized external margins ( Fig. 370 View FIGURES 362–371 ). Single female from Erzurum Province not cleared; tibia 1: 11.7.

Distribution. Known from type locality in Hatay Province, Turkey, only; specimens from Erzurum Province, Turkey, are assigned tentatively ( Fig. 446 View FIGURE 446 ).


Goteborgs Naturhistoriska Museet


Zoologisches Forschungsmuseum Alexander Koenig














Hoplopholcus atik

Huber, Bernhard A. 2020

Pholcus opilionoides

Roewer, C. F. 1959: 10