Rufocumbre, Dolibaina & Mielke & Casagrande, 2017
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publication ID |
https://doi.org/ 10.11646/zootaxa.4365.2.5 |
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publication LSID |
lsid:zoobank.org:pub:D9EA4014-FE5B-4F23-86F8-9E2853A312C2 |
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persistent identifier |
https://treatment.plazi.org/id/C3BF8E1F-F661-4C45-8F2B-B705E5BFA1D2 |
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taxon LSID |
lsid:zoobank.org:act:C3BF8E1F-F661-4C45-8F2B-B705E5BFA1D2 |
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treatment provided by |
Plazi |
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scientific name |
Rufocumbre |
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gen. nov. |
Rufocumbre gen. nov.
urn:lsid:zoobank.org:act:C3BF8E1F-F661-4C45-8F2B-B705E5BFA1D2
Type species. Rufocumbre celioi sp. nov.
Diagnosis. Rufocumbre is distinguished from all genera belonging to Moncini by the combination of the following characters: 1) eyes red; 2) antennae reaching the forewing apical hyaline spots; 3) nudum of 13 segments, in all apiculus and in part of the club; 4) labial palpus slender, third segment conical and short; 5) forewing with two groups of hyaline spots in both sexes, one apical with three small and aligned spots in R3-R4, R4- R5 and R5-M1 and the other median with two wide and somewhat rectangular spots, the distal in M3-CuA1, the proximal in CuA1-CuA2; 6) dorsal forewing of males with a triangular brand at the origin of CuA2; 7) dorsal forewing with the costal margin slightly rufous and a small cream spot over 2A; 8) ventral forewing costal area rufous to yellowish from base to apex, and a purple marginal patch from R5 to CuA1; 9) ventral hind wing costal margin rufous; 10) ventral hind wing with a dark red discal band, surrounded proximally and distally by orange to yellow bands; 11) ventral hind wing marginal patch purple, between Rs-CuA2; 12) ventral tegumen arm inserted in the most anterior portion of the tegumen; 13) anterior projection of saccus slightly longer than tegumen+uncus; 14) uncus bifid; 15) gnathos bifid, each arm with a ventral and another dorsal sclerotized plate surrounding a membranous central area; 16) fultura inferior with a short anterior projection; 17) ampulla developed in a spoonlike distal structure, with the distal margin serrated and curved inward; 18) harpe somewhat rectangular, with a robust distal spine-like projection; 19) aedeagus slender, as long as or longer than valva; 20) aedeagus distal end bifid; 21) simple scale-like cornutus, with a single spine at the middle; 22) lamella postvaginalis distally bifid; 23) ostium bursae at the base of sterigma; 24) tergum VIII with a complete spiracular opening; 25) bursa copulatrix less than four times longer than sterigma; 26) ductus bursae sclerotized; 27) corpus bursae composed by a thin membrane, with two lateral lines of signa.
Description. Head: Eye red. Labial palpus dorsally grayish, ventrally cream, anterior half of the second segment rufous brown in lateral view; first segment short, as long as wide; second segment quadrate (inner edge equal to transverse width); third segment conical and short, as long as 1/3 of the second segment length. Antennae about 60% of the forewing length; dorsally brown, ventrally with light yellow spots; nudum reddish, 13 segments, including all apiculus and part of the club.
Thorax: Dorsally brown, ventrally grayish to pale yellow; patagia and tegula rufous to brown; legs externally rufous, internally light yellow; mesotibia spined, with a distal pair of spurs; metatibia spined, with two pairs of spurs, proximal at midlength of metatibia, both meso and metatibiae with the outer spur shorter than inner.
Forewing: length 13.8-17.8 mm in males, 14.2-18.2 mm in females; apex pointed; outer margin slightly convex; tornus rounded; discal cell longer than half of the forewing length, partially opened due to the absence of dcm; 3A short; male with a sagitated brand over the origin of CuA2, superior projection longer than inferior.
Forewing dorsal: Ground color brown; two groups of hyaline spots, one subapical with three small spots in R3- M1, all aligned with each other and inclined toward the middle of the outer margin, the second group discal, with two more developed and rectangular spots, one distal in M3-CuA1, after the end of the discal cell, other proximal, in CuA1-CuA2, longer, wider and aligned with the origin of CuA1; basal third of the costal margin rufous to brown; small ellipsoid cream spot over the center of 2A; fringe light brown.
Hind wing: Oval-shaped; apex and outer margin convex; tornus rounded and projected; anal margin slightly convex; discal cell partially opened; M2 weakly marked to absent.
Hind wing dorsal: Ground color brown, base and costal areas darker; fringe light brown.
Forewing ventral: Ground color and hyaline spots as in dorsal view; costal and apical areas rufous to yellowish; marginal band purple, extending between R5-CuA1, wider in M1; fringe light brown.
Hind wing ventral: Ground color brown from inferior half of CuA2-2A to 3A; costal, submarginal and anal areas rufous; basal and postdiscal areas orange to yellow; discal band wide, dark rufous, extending from Sc+R1 to superior half of CuA2-2A, with irregular margins; marginal band purple, between Rs-CuA2, wider in M3; fringe light brown.
Abdomen: Dorsally brown. Ventrally light cream; with a central dark brown line, sometimes weakly marked.
Male genitalia: Tegumen rectangular, about 1.2 times longer than wide. Ventral tegumen arm inserted in the most anterior portion of tegumen, and longer than dorsal projection of saccus. Anterior projection of saccus triangular, longer than tegumen+uncus. Fenestra semicircular. Uncus slightly or deeply bifid; base narrower than tegumen. Gnathos completely bifid, each arm with a ventral and another dorsal sclerotized plate surrounding a membranous central area. Fultura inferior U-shaped, with a short anterior projection. Valva about two times longer than wide; costa narrow; sacculus rectangular, with 1/4 to 1/3 of the width of valva; ampulla spoon-shaped, with 1/ 3 to 1/2 of the length of valva, distal margin serrated and slightly curved inward; harpe longer than wide, about 1/3 of the width of valva, projecting a distal robust spine near the inferior margin. Aedeagus slender, as long as or longer than valva, aedeagus distal margin bifid; insertion of manica at 2/5 of the length of aedeagus, area proximal to insertion of the manica heavily sclerotized; cornutus as a simple scale-like structure, with a simple spine at the middle.
Female genitalia: Lamella antevaginalis bifid, when present. Lamella postvaginalis broad, longer than wide in ventral view; distal margin bifid; base with two short, ribbed, heavily sclerotized and ventrally curved plates. Ostium bursae at base of sterigma, with the area above weakly sclerotized. Ductus bursae sclerotized, as long as 1/ 2 to 1/3 of the length of corpus bursae. Corpus bursae ovoid, composed by thin membrane, with two lateral areas of signa, each one divided by a smooth central area. Tergum VIII divided, with a complete spiracular opening at the middle. Papilla analis almost quadrate; apophysis posterior narrow, as long as papilla analis.
Etymology. The name is a reference to the external similarity with the species belonging to the genus Cumbre , where some species of Rufocumbre were historically misplaced, in addition to the rufous aspect present in the eyes and part of the wings of its species.
Key to species of Rufocumbre
1. Uncus slightly bifid ( Figs 33–34 View FIGURES 33–34 ); lamella antevaginalis absent ( Figs 38–39 View FIGURES 38–39 )...................................... 2
– Uncus deeply bifid ( Figs 35–37 View FIGURES 35–37 ); lamella antevaginalis present ( Figs 40–42 View FIGURES 40–42 )...................................... 3
2. Uncus distal arms rounded; distal spine of harpe long, exceeding the distal margin of ampulla ( Fig. 33 View FIGURES 33–34 ); distal projections of
lamella postvaginalis short, more pointed and distant from each other ( Fig. 38 View FIGURES 38–39 )......................... celioi sp. nov.
– Uncus distal arms pointed; distal spine of harpe short, not exceeding the distal margin of ampulla ( Fig. 34 View FIGURES 33–34 ); distal projections of lamella postvaginalis longer, rounded and close to each other ( Fig. 39 View FIGURES 38–39 )............................. emeryi sp. nov.
3. Ventral hind wing basal and postdiscal areas orange ( Figs 10, 12, 14, 16 View FIGURES 1–20 ); distal arms of uncus parallel ( Figs 35–36 View FIGURES 35–37 )...... 4
– Ventral hind wing basal and postdiscal areas yellow ( Figs 18, 20 View FIGURES 1–20 ); arms of uncus divergent ( Fig. 37 View FIGURES 35–37 )......... lucasi sp. nov.
4. Ventral hind wing orange basal area with purple scales ( Figs 10, 12 View FIGURES 1–20 ); arms of uncus wide ( Fig. 35 View FIGURES 35–37 ); distal projection of lamella antevaginalis large, without a central membranous patch ( Fig. 40 View FIGURES 40–42 )........................ eberti stat. nov., comb. nov.
– Ventral hind wing orange basal area without purple scales ( Figs 14, 16 View FIGURES 1–20 ); arms of uncus narrow ( Fig. 36 View FIGURES 35–37 ); distal projection of lamella antevaginalis narrow, with a central membranous patch ( Fig. 41 View FIGURES 40–42 )........................... schneideri sp. nov.
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