Paduniella flinti, Johanson, Kjell Arne & Oláh, János, 2010

Johanson, Kjell Arne & Oláh, János, 2010, Descriptions of five new species of Paduniella from Madagascar (Trichoptera: Psychomyiidae), Zootaxa 2338, pp. 54-62 : 55-56

Paduniella flinti , new species

Figs 1–3

This species resembles P. capensis and P. s o n a in having genitalia with median branch of the inferior appendages visible and apically separated from the main branch in lateral view. The superior appendages of P. flinti and P. capensis are about similarly broad in dorsal view and the main branch of their inferior appendages curve dorsad towards apex. Paduniella flinti differs from P. c a p e n s i s and P. sona by the presence of a large ventral branch of the inferior appendages, and from P. sona in the rectangular apex of the main branch of the inferior appendages.

Male genitalia ( Figs 1–3 ). Sternite IX narrowly club-shaped in lateral view; with both ventral arms with almost straight posterior margin and convex anterior margin, 3 times longer than wide, about 0.1 mm long; with minute swelling at posterior apex; both dorsal arms narrow, needle-shaped, straight, oriented anterad, joined with tergite IX at dorsal part of genitalia, about 0.2 mm long. Ventral and dorsal arms forming narrow angle. Tergite IX long, smooth, in lateral view basally parallel-sided, reaching halfway along superior appendages; dorsal margin nearly straight. Superior appendages separate from tergite IX by longitudinal suture; parallel-sided along entire length, slightly narrower than widest part of sternite IX; weakly bending ventrad at mid-length; setae apparently restricted to distal half; apex rounded. Median branch of inferior appendages with undulated dorsal margin; about as wide as main branch of inferior appendages in lateral view; width equal in ventral and lateral views; originates before mid-length of main branch; margins nearly parallel in lateral view; covered by setae; nearly 0.1 mm long; apex truncate. Ventral lobe of inferior appendages oriented posterad; with nearly straight ventral margin in lateral view; forming pair of nearly cylindrical processes covered by setae in ventral view. Main branch of inferior appendages about parallelsided distally of origin of median branch; about as wide as superior appendages in lateral view; slightly curving dorsad; about 0.2 mm long. Phallic apparatus connected to sternite IX and tergite IX by long, straight, irregularly shaped, dorsally narrowing sternal strips. Phallobase about 0.15 mm long; narrowing distad into lobe curved dorsad; posterior half parallel-sided in lateral view, with convex dorsal and concave ventral margin; about as wide as sternal strip. Median processes nearly 0.5 mm long, needle-shaped, curved ventrad from posterior one-third; originating from anterodorsal part of phallobase; each apex exceeding phallicata. Phallicata slender at base, straight, parallel-sided, nearly 0.3 mm long; apex wide, asymmetrical, with nearly straight ventral margin, dorsal margin strongly convex. Paramere separated from phallicata immediately posterior of phallobase; running parallel with phallicata, exceeding phallicata and median process; nearly 0.4 mm long.

Holotype male, Prov. Fianarantsoa, 7 km W Ranomafana, 1100 m, 22–31.x.1988 [W. E. Steiner] ( NMNH, pinned).

Paratypes: Same data as holotype, 4 males. Same data as holotype, except 8–21.x.1988, 8 males, 2 females ( NMNH, pinned), 1 male ( NHRS, pinned), 6 males, 1 female ( NMNH, alcohol). Same data as holotype, except 600 m, 21.x.1988 [W. E. Steiner & R. van Epps], 8 females ( NMNH, pinned), 2 females ( NHRS, pinned). Same data as holotype, except 22–31.x.1988, 8 males, 2 females ( NMNH, pinned), 6 males, 1 female ( NMNH, alcohol), 2 males ( NHRS, alcohol). Same data as holotype, except 1–7.xi.1988, 8 males ( NMNH, pinned), 1 male ( NHRS, pinned), 28 males, 1 female ( NMNH, alcohol). Same data as holotype, except 1000 m, 26–29.i.1990, 2 males, 1 female ( NMNH, alcohol). Same data as holotype, except 900 m, 20– 31.i.1990, 5 males ( NMNH, pinned), 1 male ( NMNH, alcohol); same data as holotype, except 900 m, 8– 13.iii.1990, 13 males ( NMNH, pinned), 6 males, 3 females ( NMNH, alcohol). Same data as holotype, except 14–19.iii.1990, 30 males, 9 females ( NMNH, alcohol). Same data as holotype, except 900 m, 20–24.iii.1990, 6 males ( NMNH, pinned), 32 males, 13 females ( NMNH, alcohol).

Distribution. Madagascar (Province Fianarantsoa).

Etymology. This species is named after Oliver S. Flint, Jr. to acknowledge his help in making the material of this species available for this work.

Figures

FIGURES 1 – 3. Paduniella flinti, new species, holotype, male genitalia: 1, lateral; 2, phallic apparatus, lateral; 3, ventral.