Eoscartoides

Lambkin, Kevin J., 2016, Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic — Part 2, Zootaxa 4092 (2), pp. 207-218 : 213

Eoscarterella Evans, 1956

Type species. Eoscarterella media Evans, 1956, by original designation.

Diagnostic characters. Tegmen approximately 10 mm long, asymmetrically lobate, apical margin at RA2 gently convex, at M strongly convex, anterior margin fairly straight; precostal carina and marginal membrane very narrow; hypocostal carina well-developed, extending along most of costal space (Figs 11,12); sculpture comprising irregular, short, longitudinally orientated rugae, particularly coarse in basal costal space, grading to a finely punctate sculpture apically ( Fig.12 ); strigil not detected in basal costal space; hyposubcostal carina short, angulately concave; clavus not preserved; primary fork of R proximal to clavus ½ length; primary fork of RA proximal to apex of clavus, at about level of CuA fork; stem of RA1 directed anteriorly, more or less in line with stem of RA, entering costal margin at about ⅔ tegmen length, simple or with 1 or 2 anterior branches; RA2 quite extensive, occupying a large area antero-apically, with 3-5 more or less pectinate branches, the middle one forked; RA2 and RP joined by a crossvein; RP simple, not up-curved apically; M1+2 simple, M3+4 deeply forked; M and CuA joined basally by a weak crossvein; apical m-cu upright; CuA1 strongly concave; CuA2 straight, much shorter than CuA1.

Notes. The retention of a well-developed hypocostal carina is unique among Australian dysmorphoptilids and is considered plesiomorphic for the family (Dmitry Shcherbakov, pers com.). The peculiar tegmen shape and surface sculpture, and the branching pattern of M (M1+2 simple, M3+ 4 forked) are considered as the apomorphies for the genus. It is similar to Eoscartoides , the Australian Permian Belmontocarta Evans (Evans 1958) and the unnamed Triassic species from Kyrgyzstan illustrated by Shcherbakov and Popov (2002, fig.184) in sharing the evenly rounded tegmen without an emargination, the early entry of RA1 into the costal margin rather than the apex and the associated extensive and antero-apically positioned RA2.

Figures

FIGURES 10 – 12. Eoscarterella media Evans: 10 – 11, holotype UQG C 1110 / 1111, composite photograph and drawing of C 1110 and small apical lunule from counterpart C 1111 (mirrored); 12, QM F 58596, composite of part and counterpart to show surface sculpture (base mirrored).