Anaderma Lévi F Lévi, 1983

Schlacher-Hoenlinger, Monika A., Pisera, Andrzej & Hooper, John N. A., 2005, Deep-sea “ lithistid ” assemblages from the Norfolk Ridge (New Caledonia), with description of seven new species and a new genus (Porifera, Demospongiae), Zoosystema 27 (4), pp. 649-698 : 679-681

Genus Anaderma Lévi F Lévi, 1983

Anaderma Lévi F Lévi, 1983: 107 , figs 1-4.

TYPE SPECIES. — Anaderma rancureli Lévi F Lévi, 1983 by monotypy.

DEFINITION. — Anatriaenes among ectosomal spicules; microscleres include styles and amphiasters (Pisera F Lévi 2002c).

Anaderma rancureli Lévi F Lévi, 1983 ( Figs 3E ; 16 ; 30 )

Anaderma rancureli Lévi F Lévi, 1983: 107 , pl. 3: 4-5, pl. 13: 2, 3, fig. 3. — Pisera F Lévi 2 002c: 322-324, figs 1-4.

HOLOTYPE. — New Caledonian Slope ( MNHN DCL 2767 ).

MATERIAL EXAMINED (see Table 1). — Banc 1, 276- 350 m ( QM G318561, QM G318832); 305-332 m ( QM G318611); 290-378 m ( QM G318722, QM G318725); 328-340 m ( QM G318821). — Jumeau- West, 302-325 m ( QM G318781).

DESCRIPTION

Growth form

Massive conical to subspherical sponges with a simple base and a flattened upper surface with or without a shallow depression at the top. The examined specimens are about 3 to 6 cm high and 3 to 6 cm wide at the top.

Colour

Beige in etoh.

Oscules

Dispersed oscules on the upper surface, 2 mm in diameter.

Texture

Hard.

Surface ornamentation

Velvety.

Ectosomal skeleton

Outer surface of the sponge is covered by 1-3 mm thick layer without desmas which is composed of densely packed radial bunches of long oxeas and anatriaenes, which are deeply rooted in the choanosome. Sparsely distributed dichotriaenes and numerous microscleres occur in the dermal region. Microscleres also occur in great numbers in the subdermal region, which is characterised by large interconnected spaces and rare desmas.

Choanosomal skeleton

Choanosomal skeleton consists of megaclone desmas with long zygomes. The zygoses are rather weak and therefore produce a delicate skeleton. Apart from desmas there are numerous radially oriented, long oxeas.

Megascleres

Megaclone desmas smooth, slender, and poorly branched, with long zygomes and weak zygoses: 1000-1500 µm/70-100 µm.

Oxeas: 2.29 mm × 23.9 µm.

Dichotriaenes: 1.34 mm long × 741 µm (diameter of the cladome).

Anatriaenes: 91-104 µm long × 738-775 µm.

Microscleres

Microxeas (spinose, curved): 141-157 × 6.27-7.24 µm. Streptasters/amphiasters (pointed to blunt): 12.3- 14.4 × 8.36-10.2 µm.

Microstyles (spinose): 105-120 × 10-15 µm.

REMARKS

This new material conforms closely in shape and spicule geometry with material already described from the region.

Family THEONELLIDAE Lendenfeld, 1903 DEFINITION. — Polymorphic lithistids with tetraclone choanosomal desmas and phyllo- to discotriaenes as ectosomal spicules; microscleres are spinose microrhabds and/or microxeas, and streptasters and/or pseudospherasters (Pisera F Lévi 2002d).

Figures

FIG. 3. — A, Isabella mirabilis n. gen., n. sp.; B, Neoschrammeniella moreti (Lévi & Lévi, 1988); C, N. norfolkii n. sp.; D, N. castrum n. sp.; E, Anaderma rancureli Lévi & Lévi, 1983. Scale bars: 1 cm.

FIG. 16. — Anaderma rancureli Lévi & Lévi, 1983; A, B, outer surface of the choanosomal skeleton; note long spines of megaclones; C, D, details of articulation of megaclones; E, F, smooth megaclone choanosomal desma. Scale bars: A-C, E, F, 500 µm; D, 100 µm.

FIG. 30. — Anaderma rancureli Lévi & Lévi, 1983; A, B, section perpendicular to the outer surface showing subectosomal spaces and bunches of anatriaenes and oxeas in fan-like organisation; C, microxeas; D, styles; E, ectosomal anatriaene; F, streptaster/amphi- aster. Scale bars: A, B, 1 µm; C, 20 µm; D, 10 µm; E, 50 µm; F, 5 µm.