Drassodes kaszabi Loksa, 1965

Fomichev, A. A. & Marusik, Y. M., 2015, First Description Of The Male Of Drassodes Kaszabi (Aranei, Gnaphosidae), Vestnik Zoologii 49 (5), pp. 467-470 : 467-470

Drassodes kaszabi Loksa, 1965 ( fig. 1–11 )

D. kaszabi Loksa, 1965: 27 , fig. 46 (♀); Marusik, Logunov, 1995: 183, fig. 11–12 (♀).

M a t e r i a l e x a m i n e d. Mongolia: Bayan-Ölgii Aimag: Khongor-Olongiyn-Gol River Valley, 34 km

ENE from Tolbo Village , 48°31ʹ N, 90°43ʹ E, mountain stony steppe with rocks, 2300 m, 5.05.2012, 1 ♀ ; Khovd Aimag: Arshantyn-Nuruu Mt. Range , 46°21ʹ N, 91°15ʹ E, mountain stony steppe with rocks, 1700–2100 m, 15.05.2012, 2 Ơ, 2 ♀ ; Mongolian Altai Mt. Range (S slope), Bodonchiyn-Gol River Basin, Khondiyn-Gol River Valley , 46°07ʹ N, 92°30ʹ E, semi-desert steppe with rocks, at night, 1800 m, 17.05.2012, 1 ♀ ; foothills of Muchar- Uul Mt. , 47°09ʹ N, 92°11ʹ E, mountain stony steppe with rocks, 1900–2000 m, 5– 6.05.2012, 1 Ơ, 2 ♀ ( AF) ( ISEA) .

D i a g n o s i s. Males of D. kaszabi are similar to those of D. pubescens (Thorell, 1876) , D. dispulsoides Schenkel, 1963 and D. cupa Tuneva, 2005 in having a modified median apophysis (transverse, with bifurcated tip) and a straight embolus, differing from these two species by having a long tibial apophysis, almost as long as the palpal tibia (2 or more times shorter in similar species). The tibial apophysis of D. cupa is sharply pointed, but it is not in D. kaszabi . Females of D. kaszabi differ from other species by having a large epigynal fovea with distinct margins and a small median plate (two times shorter than fovea height).

D e s c r i p t i o n. Male. Total length 10.3. Carapace: 4.8 long, 3.4 wide. Colouration. Carapace, sternum and legs yellow-brown ( fig. 1 ). Chelicerae brown, with 3 promarginal teeth and 1 small retromarginal tooth ( fig. 2 ). Maxillae and labium light-brown. Abdomen beige, without pattern.

Leg measurements:

Leg Fe Pt Ti Mt Ta Total I 4.1 2.2 3.8 3.25 2.05 15.4 II 3.85 2.1 3.25 2.95 1.95 14.1 III 3.65 1.75 2.8 2.95 1.9 13.05 IV 4.5 2.0 3.8 4.7 2.25 17.25

Leg spination:

Leg Fm Ti Mt

I d1–1–0 p0–1–1 p1–0–1 v2–0–2 v2–0–0

II d1–1–0 p0–1–1 p1–0–1 v2–0–2 v2–0–0

III d1–1–1 p1–1–1 r0–1–1 d1–0–0 p1–1–1 r2–0–1 v2–2–2 d1–2–0 p1–1–0 r1–1–0 v2–2–0 IV d1–1–1 p0–1–1 r0–2–1 d1–0–1 p1–1–1 r2–0–1 v2–2–2 d1–2–0 p1–1–0 r1–1–0 v2–0–2

Palp as in fig. 3–8 . Tibia with 2–3 dorsal and 2 prolateral spines. Tibial apophysis long, almost as long as tibia. Spermophor (Sp) with three loops, terminal part directed downward (basally). Median apophysis (Ma) modified, without hook, much wider than long, almost as wide as tegulum, tip bifurcated. Conductor (Co) large, membranous with serrated margins. Embolus strong and straight, slightly shorter than median apophysis width.

F e m a l e. See Loksa (1965) and Marusik, Logunov (1995) for description. Total length 14.0. Carapace: 4.8 long, 3.3 wide. Colouration as in male.

Leg measurements:

Leg Fe Pt Ti Mt Ta Total I 3.85 2.1 3.15 2.65 1.8 13.55 II 3.65 1.9 2.9 2.55 1.75 12.75 III 3.5 1.75 2.75 2.85 1.8 12.65 IV 4.5 2.0 3.8 4.65 2.0 16.95

Leg spination:

Leg Fm Ti Mt

I d1–1–0 p0–1–1 v2–2–0 v2–0–0

II d1–1–0 p0–1–1 p0–0–1 v2–2–0 v2–0–0

III d1–1–1 p0–1–1 d1–0–0 p1–1–1 r2–0–1 v1–2–2 d1–2–0 p1–1–0 r1–1–0 v2–2–0

IV d1–1–1 p0–1–1 r0–1–1 d1–0–1 p1–1–1 r2–0–1 v2–2–2 d1–2–0 p1–1–0 r1–1–0 v2–0–2

Epigyne as in fig. 9–11 . Fovea with distinct margins, 1.6 times wider than high. Median plate (Mp) about 2.2 times thinner than cavity and almost 2 times shorter than cavity. Each receptacle with 2 large chambers; receptacles span a distance 0.77 times less than cavity width.

S i z e v a r i a t i o n. Male body length varies from 10.0 to 10.7, carapace 4.1–4.8 long and 3.0–3.4 wide. Female body length varies from 9.6 to 14.0, carapace 4.5–5.6 long and 3.2–3.95 wide.

D i s t r i b u t i o n. Western Mongolia (Bayan-Ölgii, Khovd, Uvs, Bayankhongor and Övörkhangai Aimags) and Western Tuva ( Russia).

We thank R. V. Yakovlev (Barnaul, Russia), A. N. Nakonechnyi (Novosibirsk, Russia), A. I. Shmakov (Barnaul, Russia), G. G. Khabiev (Aktash, Russia) and U. Beket (Ölgii, Mongolia) for organizing and undertaking an expedition to Mongolia, from where the material treated in this paper was collected. Thanks to S. Koponen (University of Turku, Finland) and R. Yu. Dudko (ISEA) for providing institutional facilities. English of the earlier draft was kindly checked and corrected by Sarah Crews (California Academy of Sciences, USA).

Figures

Fig. 1–11. Diagnostic characters of Drassodes kaszabi: 1 — male habitus, dorsal; 2 — male chelicera, posterior; 3–5 — male palp, ventral, retro- and prolateral; 6–8 — bulbus, dorso-retrolateral, prolateral and ventral; 9–10 — epigyne, ventral; 11 — epigyne, dorsal. Scale bar 0.2 mm if not otherwise indicated. Abbreviations: Co — conductor; Ma — median apophysis; Mp — median plate; Sp — spermophor.

Fig. 12. Distribution records of Drassodes kaszabi:  literature data;  new records.