Draculoides gnophicola ( Harvey, Berry, Edward and Humphreys, 2008 )

Abrams, Kym M., Huey, Joel A., Hillyer, Mia J., Didham, Raphael K. & Harvey, Mark S., 2020, A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara, Zootaxa 4864 (1), pp. 1-75 : 45-47

publication ID

https://doi.org/10.11646/zootaxa.4864.1.1

publication LSID

lsid:zoobank.org:pub:A5F51A7F-83DA-4C77-A85C-0FCF8A400CF2

DOI

http://doi.org/10.5281/zenodo.4417335

persistent identifier

https://treatment.plazi.org/id/03CE87D9-FFAA-FFFD-CC8B-0494FF38FCB5

treatment provided by

Plazi

scientific name

Draculoides gnophicola ( Harvey, Berry, Edward and Humphreys, 2008 )
status

 

Draculoides gnophicola ( Harvey, Berry, Edward and Humphreys, 2008)  

( Figs. 1–8 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 , 22–23 View FIGURE 22 View FIGURE 23 )

http://zoobank.org/NomenclaturalActs/ C8597E44-F7BC-43AA-9DA1-F2D7B9B423DF

Paradraculoides gnophicola Harvey, Berry, Edward and Humphreys 2008:188–190   , figs. 43–46.

Draculoides gnophicola (Harvey, Berry, Edward and Humphreys)   : Abrams et al. 2019 MPE 106532: 8, fig. 2.

Material examined. Holotype female. AUSTRALIA: Western Australia: Mesa G (Borehole 0159A), 22.4 km SW. of Pannawonica , 21°44’20”S, 116°08’06”E, March–April 2008, M. Greenham, D. Kamien and L. Mould ( WAM T 65763 View Materials ). GoogleMaps  

Paratypes. AUSTRALIA: Western Australia: 1 ♀, Mesa G (Borehole 0014A), 22.4 km SW. of Pannawonica , 21°43’57”S, 116°07’01”E, 21 December 2004, G. Humphreys, M. Greenham ( WAM T 63312 View Materials ) GoogleMaps   ; 1 ♀, Mesa G (Bore- hole 0057A), 21 December 2004, G. Humphreys, M. Greenham ( WAM T 63316 View Materials )   .

Other material. AUSTRALIA: Western Australia: 1 ♂, Mesa G, 23.9 km SW. of Pannawonica , 21°43’57”S, 116°07’00”E, 21 October 2009, troglofauna trap, J. Alexander, E. Harris (Biota Environmental Sciences, MEG- DC014AP1 T3-4 ) ( WAM T 99563 View Materials ) (DNA: COI); see Abrams et al. (2019) GoogleMaps   .

Diagnosis. The shape of the male flagellum of Draculoides gnophicola   most closely resembles D. affinis   , D. anachoretus   , D. bythius   , D. cochranus   , D. eremius   and D. kryptus   , especially in the presence of a broad base. It differs from D. affinis   by the position of dm4 level with dl3 (dm4 is slightly anterior to dl 3 in D. affinis   ) and vm1 which is slightly posterior to vm2 (vm1 more posterior to vm 2 in D. affinis   ), from D. anachoretus   and D. cochranus   by the posterior position of dm4 (anterior in D. anachoretus   and D. cochranus   ), level with dl3; from D. bythius   in the close proximity of dl1 and vl1 (far apart in D. bythius   ); from D. kryptus   by the sub-distal position of vl2 (on distal margin in D. kryptus   ), and from D. eremius   by the raised antero-dorsal region of the flagellum, which is flat in D. eremius   . Females of this species differ from all others in the position of flagellar seta dm4, which is much closer to dl1 than any other Draculoides   . Draculoides gnophicola   can be diagnosed from all other Draculoides   species that were sequenced at COI and 12S by the 50bp mini-barcodes shown in Figures 3 View FIGURE 3 and 5 View FIGURE 5 . Draculoides gnophicola   can be diagnosed from all other Draculoides   species that were sequenced at ITS2 ( Fig. 7 View FIGURE 7 ) except for D. anachoretus   , D. bythius   , D. eremius   , D. kryptus   , D. mckechnieorum   , D. warramboo   , D. immortalis   , D. belalugosii   , D. christopherleei   , D. piscivultus   and D. akashae   , which are not distinguishable using the ITS2 mini-barcode.

Description. Adult male (WAM T99563 View Materials )

Colour. Yellow-brown; propeltidium and pedipalps somewhat darker.

Cephalothorax. Propeltidium with 2+1 apical setae in a triangular formation on anterior process and 2 + 2 + 1 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 14 setae (including 2 sternapophysial setae); posterior sternum triangular with 6 setae.

Chelicera. Fixed finger with 2 large teeth plus 5 smaller teeth between these including 1 tiny lateral tooth on proximal tooth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1);. G2 composed of 7 setae; G3 composed of 3 setae; internal face of chelicera with 3 short whip-like setae (G4); brush at base of fixed finger composed of 8 setae (G5A), each densely pilose in distal half and G5B composed of 11 setae; G6 and G7 composed of one seta. Movable finger serrula composed of ~18 long lamellae, blunt guard tooth present subdistally; 1 large accessory tooth present.

Pedipalp. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 8 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.33 × length of tarsus.

Legs. Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.

Abdomen. Chaetotaxy of tergite I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 2; segment XII with small dorsal process (♂ only).

Flagellum. Male: Dorsoventrally compressed ( Figs. 22 View FIGURE 22 D–F, 23A–C); 1.9 × longer than broad; seta dm1 situated dorso-medially, closer to anterior margin; seta dm4 close to posterior margin, level with dl3; dl1posterior to vl1; dl3 on posterior margin; vm2 situated slightly anterior to vm1; vm5 situated midway between vl2 and vm3; left vm3 slightly posterior to right vm3; one pair of microsetae dorsally, slightly posterior to dm1, six pairs of microsetae between vl1 and dl3.

Dimensions (mm). Male (WAM T99563 View Materials ): Body length 3.17. Propeltidium 1.15/0.62. Chelicera 0.73. Flagellum 0.37/0.19. Pedipalp: trochanter 0.50, femur 0.48, patella 0.60, tibia 0.58, tarsus 0.29, claw 0.10.

Remarks. Harvey et al. (2008) named this species from several females and juveniles collected from Mesa G ( Fig. 1B View FIGURE 1 ). Additional specimens, including a male that is described here, have since been collected from the same mesa.

WAM

Western Australian Museum

T

Tavera, Department of Geology and Geophysics

COI

University of Coimbra Botany Department

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Schizomida

Family

Hubbardiidae

Genus

Draculoides

Loc

Draculoides gnophicola ( Harvey, Berry, Edward and Humphreys, 2008 )

Abrams, Kym M., Huey, Joel A., Hillyer, Mia J., Didham, Raphael K. & Harvey, Mark S. 2020
2020
Loc

Paradraculoides gnophicola

Harvey, Berry, Edward and Humphreys 2008: 188 - 190
2008