Chasmatonotoides deharvengi, Published, 2007

Published, First, 2007, Chironomid midges from early Eocene amber of France (Diptera: Chironomidae), Zootaxa 1404, pp. 1-66: 35-36

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Chasmatonotoides deharvengi

n. sp.

Chasmatonotoides deharvengi   n. sp.

( Figs. 32, 33)

Etymology: This species is named for Dr. Louis Deharveng, entomologist at the MNHN.

Diagnosis: Body ca. 2.5 mm long; 1 prealar seta; squama bare; gonostylus covered with numerous short setae.

Description: Head 0.5 mm long, deformed; ocelli absent; antenna much longer than head, distinctly hairy, with 13 flagellomeres covered with long setae, pedicel broad and short, rounded, 13th flagellomere very long; eye bare, with small dorsomedial extension, with 1 row of ommatidia at minimum width; mouthparts lacking functional mandibles; frontal setae absent, numerous postocular and inner and outer vertical setae. Thorax 0.78 mm long, 0.36 mm wide, 0.76 mm high; postnotum bare, without longitudinal median groove; surface of scutellum with 1 long seta visible; scutal tubercle absent; acrostichals absent, dorsocentrals uniserial; scutum with median longitudinal groove; 1 prealar seta, no supraalar setae; epimeron II and preepisternum bare. Wing macropterous, 1.52 mm long, 0.36 mm wide, hyaline, membrane bare, but with coarse punctation; anal vein An 2 absent; radius with 3 branches R 1, R 2+3 and R 4+5, R 2+3 weakly sclerotized not divided into R 2 and R 3 and nearly to R 4+5; only M 1+2 and M 3+4 present; cross­vein MCu absent; squama bare. Halter 0.2 mm long. Fore femur 0.5 mm long, tibia 0.54 mm long, tarsus 1.2 mm long; mid femur 0.56 mm long, tibia 0.54 mm long; hind femur 0.54 mm long, tibia 0.58 mm long, tarsus 0.88 mm long; ta4 of all legs cylindrical, not cordiform; fore tibia with 1 long spur, mid and hind tibiae with 2 long spurs, hind tibial comb well developed. Abdomen 1.72 mm long; gonostylus 0.05 mm long, 0.03 mm wide, simple, covered with numerous setae; gonocoxite 0.1 mm long, 0.05 mm wide, with numerous long setae on outer surface; anal point narrow and elongate, 0.15 mm long, 0.1 mm wide at base, apex seemingly truncated.

FIGURE 33. Chasmatonotoides deharvengi   n. sp., holotype PA 9563, drawing of male genitalia (scale bar = 0.1 mm).

Discussion: The body of C. deharvengi   n. sp. (2.5 mm long) is distinctly longer than that of C. oesiensis   (1.7 mm long). It has one prealar seta instead of two, as in C. oesiensis   . For these reasons, we recognize a new species. Material: Holotype PA 9563, (male).