Tokunagaia azari, Published, 2007

Published, First, 2007, Chironomid midges from early Eocene amber of France (Diptera: Chironomidae), Zootaxa 1404, pp. 1-66: 43-44

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Tokunagaia azari

n. sp.

Tokunagaia azari   n. sp.

( Figs. 38 ­40)

Etymology: We name this species for our colleague Dr. Dany Azar (University of Lebanon).

FIGURE 39. Tokunagaia azari   n. sp., holotype PA 11435, photograph of male genitalia.

FIGURE 40. Tokunagaia azari   n. sp., holotype PA 11435, drawing of male genitalia (scale bar = 0.1 mm).

Diagnosis: Eye with short wedge­shaped dorsomedial extension; antennal ratio 0.46; wing membrane without macrotrichia; anal point absent; gonostylus simple, with low crista dorsalis, tergite IX with less than 20 setae.

Description: Head 0.25 mm long, deformed; ocelli absent; antenna 0.41 mm long, much longer than head, distinctly hairy, with 13 flagellomeres covered with long setae (shortest 0.01 mm long, longest 0.19 mm long), pedicel broad and short, rounded, 13 th flagellomere 0.15 mm long; antennal ratio 0.46; eye bare, with small dorsomedial extension, with 2 rows of ommatidia at minimum width; mouthparts lacking functional mandibles; 5 palpomeres with setae; 1 postocular; frontal and inner and outer vertical setae not visible, probably absent.

Thorax 0.34 mm long, 0.19 mm wide, 1.21 mm high; postnotum bare, with longitudinal median groove; surface of scutellum bare; without scutal tubercle; acrostichals absent, dorsocentrals uniserial; scutum without median longitudinal groove; 2 prealar setae, no supraalar setae; epimeron II and preepisternum bare. Wing macropterous, 0.88 mm long, 0.2 mm wide, hyaline, membrane bare, but with coarse punctation; anal vein An 2 absent; radius with 3 branches R 1, R and R 4+5, R 2+3 not divided into R 2 and R 3; only M 1+2 and M 3+4 present; cross­vein MCu absent; squama with 2 setae. Halter 0.1 mm long. Fore femur 0.18 mm long, tibia 0.33 mm long, tarsus 0.43 mm long; mid femur 0.2 mm long, tibia 0.26 mm long, tarsus 0.28 mm long; hind femur 0.16 mm long, tibia 0.29 mm long, tarsus 0.38 mm long; ta4 of all legs cylindrical, not cordiform; fore tibia with 1 long spur, mid and hind tibiae with 2 long spurs, hind tibial comb well developed; small pulvilli present. Abdomen 0.82 mm long; tergite IX with about 5­10 setae; gonostylus 0.04 mm long, 0.008 mm wide, simple, elongate, with low crista dorsalis, hinged to gonocoxite and folded inward; gonocoxite 0.05 mm long, 0.03 mm wide, with long setae apically and on outer surface; anal point absent.


Discussion: In the key to Holarctic orthocladiine genera of Cranston et al. (1989), this Orthocladiinae   falls in the genus Tokunagaia   because of the following characters: wing well developed; without thick clavus; scutum without median longitudinal groove; eye bare; wing membrane without macrotrichia; epimeron II and preepisternum bare; no lateral spines on spurs of mid and hind tibiae; anal point without strong lamellate setae laterally and apicodorsally; squama with at least 1 seta; small pulvilli, never as long as half claw length; hind tibial comb well developed; tergite IX without strongly elevated ridge; anal point absent; gonostylus simple; and eye with short wedge­shaped dorsomedial extension. The species is near the group of extant species including T. scutellata   , T. rowensis   , and T. rectangularis   because it has no anal point, a gonostylus with a low crista dorsalis, antennal ratio 0.46, and tergite IX with fewer than 20 setae ( Halvorsen & Saether 1987).

Material: Holotype PA 11435, (male).