Prodotia, Dall, 1924

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 53-54

Prodotia ? wesselyi nov. sp.

Figs 27A–C

Jania angulosa Brocc. —Hoernes & Auinger 1885: 231, pl. 27, figs 13–14 [non Janiopsis angulosa ( Brocchi, 1814) ].

Type material. Holotype: NHMW 1865 /0001/1085, SL: 44.0 mm, MD: 19.3 mm, Grund ( Austria), illustrated in Hoernes & Auinger (1885: pl. 27, fig. 13), Figs 27A 1 –A 2 . Paratypes: NHMW 1869 /0001/0274, SL: 46.6 mm , MD: 21.6 mm, Grund ( Austria), illustrated in Hoernes & Auinger (1885: pl. 27, fig. 14), Figs 27B 1 –B 2 . NHMW 1855 /0045/0689, SL: 43.8 mm , MD: 19.7 mm, Grund ( Austria), Figs 27C 1 –C 2 .

Type locality. Grund ( Austria), North Alpine-Carpathian Foreland Basin .

Type stratum. Sandy silt of the Grund Formation.

Age. Middle Miocene, early Badenian (Langhian).

Etymology. In honor of Godfrid Wessely (OMV-AG), grandseigneur of the geology of the Vienna Basin.

Diagnosis. Moderately large, moderately slender fusiform shell with prominent axial ribs overrun by several close-set spiral cords; columella with two merged denticles close to angled transition to siphonal canal. Outer lip thickened by terminal varix with eight or ten elongate denticles placed some distance behind peristome.

Description. Moderately large, moderately slender fusiform shell of seven teleoconch whorls with high spire; apical angle ~50°. Spire whorls moderately convex with faintly concave subsutural ramp, weakly angled at shoulder forming periphery mid-whorl, separated by moderately incised, shallowly undulating suture. Sculpture of broad, prominent axial ribs overrun by several close-set spiral cords of alternating strength. Last whorl high, attaining 60% of total height, broad slightly concave to convex subsutural ramp delimited by weak shoulder, strongly constricted at base, bearing about 12 axial ribs, persisting over base, overrun by spirals of alternating strength over entire surface. Aperture elongate pyriform, moderately narrow. Columellar callus forming broad rim, sharply delimited from base. Columella moderately excavated with two merged denticles close to angled transition to siphonal canal. Anal canal broad U-shaped, accentuated by prominent parietal denticle. Weak anal denticle. Outer lip thickened by terminal varix with eight or ten elongate denticles placed some distance behind peristome, roughly arranged in pairs. Siphonal canal moderately long, moderately wide, slightly deflected to the left.

Discussion. This species was placed in Janiopsis Rovereto, 1889 by Hoernes & Auinger (1885) but lacks the strong anal denticle characteristic to that genus and has paired labial denticles. We provisionally place it in Prodotia based on its apertural features but note that the prominent axial ribs of the Paratethyan species are atypical for that genus.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund ( Austria) (Hoernes & Auinger 1885).

Genus Tethyspollia nov. gen.

Type species. Pollia mariae Hoernes & Auinger, 1890 . Middle Miocene , Central Paratethys Sea .

Diagnosis. Medium-sized, biconic shell with prominent sculpture of primary and secondary spiral cords overriding broad axial ribs. Aperture narrowed by apertural armature, with several columellar folds, anal canal indistinct, lacking parietal and anal denticles. Outer lip with very prominent denticles with upper three forming palatal swelling, placed some distance from adapical tip of aperture.

Description. Medium-sized, biconic shell with slightly coeloconoid to conical early spire. Early teleoconch whorls low, almost flat sided with three prominent spiral cords overriding widely spaced axial ribs. Suture deeply incised causing marked incisions in shell profile. Axial and spiral sculpture prominent on later teleoconch whorls. Last whorl attaining ~65–70% of total height, strongly convex at periphery, base moderately constricted. Aperture narrowed by apertural armature. Columella weakly to moderately excavated. Columellar callus forming prominent rim, sharply delimited from base, with three to four broad columellar folds.Anal canal indistinct. No parietal denticle, no anal denticle. Outer lip thickened with very prominent, blunt denticles, top three forming poorly delimited palatal swelling, weakening towards siphonal canal. Adapical denticle very prominent, placed some distance from adapical tip of aperture. About six long lirae deep inside aperture. Siphonal canal moderately short to moderately long.

Etymology. Referring to the Tethys Ocean and to the Pisaniidae genus Pollia J.E. Gray, 1833 .

Included species. Pollia mariae Hoernes & Auinger, 1890 and Cantharus (Pollia) beregovi Kojumdgieva in Kojumdgieva & Strachimirov, 1960.

Stratigraphic and geographic range. Middle Miocene, Langhian, Central Paratethys Sea ( Bulgaria, Romania).

Paleoenvironment. Unknown.

Discussion. Tethyspollia nov. gen. is characterized by the absence of parietal and anal denticles and thus belongs to a group of mainly Indo-West Pacific species with weak or subobsolete parietal and anal denticles. These species were placed by Fraussen & Stahlschmidt (2016a) in Clivipollia Iredale, 1929 , Falsilatirus Emerson & Moffitt, 1988 , Speccapollia Fraussen & Stahlschmidt, 2016 and Minioniella Fraussen & Stahlschmidt, 2016 . Of these genera, Tethyspollia is closest to Speccapollia [type species Ricinula recurva Reeve, 1846a ; original designation by Fraussen & Stahlschmidt (2016a: 40)]. The depressed early teleoconch whorls with three prominent spiral cords of Tethyspollia mariae are very similar to those of Speccapollia in general, and the extant Speccapollia africana Fraussen & Stahlschmidt, 2016 , from Mozambique, develops a very similar shell shape. In addition, Tethyspollia mariae develops delicate lirae deep inside the aperture, also seen in Speccapollia . Speccapollia species, however, are smaller (around 10 mm in height), the anal canal is more deeply incised, flaring the adapical tip of the outer lip, and have a very weak anal denticle which is separated from the outer lip denticles by a distinct concavity. Moreover, the denticles on the outer lip of Speccapollia are much smaller.

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 27. Prodotia? Wesselyi nov. sp. A1–A2. NHMW 1865/0001/1085, Grund (Austria). B1–B. NHMW 1869/0001/0274, Grund (Austria). C1–C2. NHMW 1855/0045/0689, Grund (Austria).