Euthria subnodosa (Hoernes & Auinger, 1890)

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 86-88

publication ID	https://doi.org/ 10.11646/zootaxa.5427.1.1
publication LSID	lsid:zoobank.org:pub:923206B0-E8C5-4FD5-B882-55009ABB0282
persistent identifier	https://treatment.plazi.org/id/03CE9F1C-FFCF-0C10-FF65-FF7AED27FD02
treatment provided by	Plazi
scientific name	Euthria subnodosa (Hoernes & Auinger, 1890)
status

Treatment

Figs 34J View FIGURE 34 , 45A–E View FIGURE 45

Fusus Bredai Michti. —Hörnes 1853: 284, pl. 31, figs 8a–b [non Aplus bredai ( Michelotti 1847) ].

Fusus Bredai Michelotti — Neugeboren 1854: 187 [non Aplus bredai ( Michelotti 1847) ].

Pollia Badensis View in CoL nobis—Hoernes & Auinger 1890: 238.

* Fusus (Euthria) subnodosus nov. form.—Hoernes & Auinger 1890: 258, pl. 32, figs 1–2.

Fusus corneus — Quenstedt 1884: 609, pl. 209, fig. 44 [non Euthria cornea ( Linnaeus, 1758) View in CoL ].

Fusus (Euthria) subnodosus R. Hoern. et Auing. — Macovei 1909: 147, pl. 11, fig. 7.

Euthria Januszkiewiczi Friedb. View in CoL — Friedberg 1912: 156, pl. 9, fig. 8.

Euthria Januszkiewiczi Friedb. View in CoL — Friedberg 1938: 138.

Euthria januszkiewiczi Friedberg, 1912 — Zelinskaya et al. 1968: 190, pl. 45, figs 12–13.

Euthria subnodosa fortestriata n. ssp. — Csepreghy-Meznerics 1969: 86, pl. 4, figs 11, 13.

Euthria subnodosa fortestriata Csepr. -Mezn.— Csepreghy-Meznerics 1972: 28, pl. 11, figs 11–12.

E [uthria]. (E [uthria].) subnodosa (R. Hörn. et Au.) — Sieber 1958: 150.

C [antharus]. (P [ollia]). badensis (R. Hörn. et Au.) — Sieber 1958: 151.

Buccinulum (Euthria) subnodosum Hoernes et Auinger , sp. 1890— Glibert 1963: 72.

Euthria (Euthria) intermedia View in CoL (Michelotti in Sowerby, 1839)— Atanacković 1985: 146, pl. 32, figs 17–18 [non Euthria intermedia ( Michelotti, 1847) View in CoL ].

Euthria subnodosa fortestriata Csepreghy-Meznerics, 1969 — Palfy et al. 2008: 106.

A [nna] badensis (Hörnes and Auinger, 1890) — Vermeij 2006: 72.

non Euthria subnodosa Hoernes & Auinger 1890 — Glibert 1952: 323, pl. 9, fig. 3 [= Euthria sp. ].

non Euthria subnodosa Hoernes et Auinger — Csepreghy-Meznerics 1969: 86, pl. 4, figs 20–21 [= Euthria odiosa nov. sp.]. non Euthria subnodosa Hoernes et Auinger — Csepreghy-Meznerics 1972: 28, pl. 11, figs 7–8 [= Euthria depressospira Bandat, 1943 ].

non Euthria subnodosa (Hoernes et Auinger) — Kovács 2018: 182, figs 13–14 [= Euthria sp. ].

Type material. Lectotype (designated herein): NHMW 1949 View Materials /0005/0023a, SL: 33.7 mm, MD: 16.0 mm, Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 1), Figs 45A View FIGURE 45 1 –A View FIGURE 1 2 View FIGURE 2 . Paralectotypes: NHMW 1855 View Materials /0045/0452, SL: 32.6 mm , MD: 15.3 mm, Gainfarn ( Austria), Figs 45B View FIGURE 45 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1949 View Materials /0005/0023b, SL: 25.9 mm , MD: 14.6 mm, Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1890: pl. 32, fig. 2), Figs 45C View FIGURE 45 1 – C View FIGURE 1 2 View FIGURE 2 . NHMW 1846 View Materials /0037/0274, SL: 27.7 mm , MD: 14.3 mm, Enzesfeld or Gainfarn ( Austria), Figs 45D View FIGURE 45 1 –D View FIGURE 1 2 View FIGURE 2 .

NHMW 2023 View Materials /0352/0002, SL: 36.9 mm , MD: 17.9 mm, Mikulov ( Czech Republic); Figs 45E View FIGURE 45 1 –E View FIGURE 1 2 View FIGURE 2 . Additional paratypes: 23 spec. , NHMW 1860 View Materials /0001/0226, Mikulov ( Czech Republic); 12 spec. , NHMW 1860 View Materials /0001/0225, Steinebrunn ( Austria); 48 spec. , NHMW 1846 View Materials /0037/0274, Steinebrunn and Gainfarn ( Austria), Fig. 34J View FIGURE 34

Additional material. 38 spec., NHMW 1846 View Materials /0037/0273, Steinebrunn , Gainfarn and Vienna / Pötzleinsdorf ( Austria) ; 10 spec., NHMW 1856 View Materials /0050/0123, Gainfarn ( Austria) ; 4 spec., NHMW 1855 View Materials /0045/0453, Gainfarn ( Austria) ; 3 spec., NHMW 1863 View Materials /0015/0681, Niederleis ( Austria) .

Revised description. Medium-sized, moderately broad, biconic shell of up to seven teleoconch whorls; apical angle 47–55°. Protoconch conical of 1.5 strongly convex whorls; diameter: 720 μm, height: 690 μm. Early teleoconch whorls with straight subsutural ramp, weakly shouldered, bearing broad, widely spaced axial ribs, overrun by five narrow spiral cords. Adapically axial ribs weaken, fading on third teleoconch whorl; spiral cords increasing in number by intercalation of secondary and tertiary spirals. Later spire whorls faintly concave to flat-sided, separated by weakly incised suture, bearing numerous low primary and secondary spiral cords; axials reduced to blurred ribs most prominent at abapical suture. Last whorl attaining ~75% of total height, with broad, weakly concave subsutural ramp, rounded at shoulder that is smooth or with about ten broad, weak axial swellings, spiral sculpture of weak, broad spiral cords, most prominent over subsutural ramp and base, base moderately constricted, fasciole indistinct. Aperture moderately wide, pyriform. Columella moderately and broadly excavated, twisted at siphonal canal. Columellar callus indistinct, not delimited from base. Anal canal narrowly incised with weak parietal denticle; no anal denticle. Outer lip thin with about ten very prominent lirae placed some distance behind peristome, extending deep within aperture; some lirae start deeper within aperture resulting in bifurcated appearance. Siphonal canal moderately long, wide, slightly deflected to the left, weakly recurved, shallowly notched.

Paratethyan synonyms. Euthria subnodosa fortestriata Csepreghy-Meznerics, 1969 , holotype: M.70.522, Gyükerfői-dombok, Balaton ( Hungary). The holotype is a specimen with very prominent spiral sculpture of close-set spiral cords. We interpret it as extreme morphotype of Euthria subnodosa and list it tentatively as synonym of Euthria subnodosa . Euthria januszkiewiczi Friedberg, 1912 (SL: 20 mm, MD: 11 mm) from Zborów ( Ukraine) is based on a small, poorly preserved specimen of Euthria subnodosa .

Pollia badensis Hoernes & Auinger, 1890 , is based on a single specimen (holotype: SL: 16 mm, MD: 9 mm) from Bad Vöslau ( Austria), illustrated in Hörnes (1853: pl. 31, fig. 8). The specimen is lost and the illustration in Hörnes (1853) was said by Hoernes & Auinger (1890) to be not well depicted. In addition, those authors noted that the last whorl bears traces of trauma, which might have influenced the shape of the last whorl. Therefore, a comparison with other species is difficult. The illustration in Hörnes (1853) fits well with Euthria subnodosa , a species, which was not known to Hörnes (1853). Pollia badensis Hoernes & Auinger, 1890 and Euthria subnodosa Hoernes & Auinger, 1890 were published in the same monograph. As first revisers we give precedence to the name Euthria subnodosa Hoernes & Auinger, 1890 . Aplus bredai ( Michelotti, 1847) , from the Tortonian of Stazzano ( Italy), with which this species was confused by Hörnes (1853), differs in its sculpture of close-set spiral cords bearing subquadrate tubercles (see Michelotti 1847: 406, pl. 10, fig. 8).

Discussion. This species is characterized by its biconic shell much reduced axial sculpture restricted to the earliest teleoconch whorls and faint tubercles at the shoulder of the last whorl in some specimens, relatively prominent spiral sculpture for the genus, although somewhat variable, and very strong lirae within the outer lip. Euthria minor Bellardi, 1873 , from Early Miocene of the Colli Torinesi, is slightly reminiscent but differs in its slenderer outline and more prominent axial ribs ( Bellardi 1873; pl. 13, fig. 24; Ferrero Mortara et al. 1981: pl. 6, fig. 13). Specimens from the Langhian of the Loire Basin, described by Glibert (1952) as Euthria subnodosa are only distantly related to the Paratethyan species and differ in their marked shoulder placed below mid-whorl and the prominent denticles on the columella.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Friedberg 1938); Polish-Ukrainian Fore-Carpathian Basin: Zborów (Zboriv) ( Ukraine) ( Friedberg 1938); North Alpine-Carpathian Foreland Basin: Hrušovany nad Jevišovkou, Lysice ( Czech Republic) (Hoernes & Auinger 1890); Vienna Basin: Enzesfeld, Gainfarn, Niederleis, Steinebrunn, Vienna /Pötzleinsdorf ( Austria) (Hoernes & Auinger 1890; hoc opus); Mikulov ( Czech Republic) (Quenstedt 1884; Hoernes & Auinger 1890); Southern Pannonian Basin: Hrvaćani, Miljevići ( Bosnia and Herzegovina) ( Atanacković 1985); Bahna Basin ( Romania) ( Macovei 1909).

MD	Museum Donaueschingen

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Tudiclidae
Genus	Euthria