Euthria dellabellai, Harzhauser & Landau, 2024

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 69-70

Euthria dellabellai nov. sp.

Figs 36A–D

Fusus Puschi Andr. —Hörnes 1853: 282 (pars) [non Euthria puschii ( Andrzejowski, 1830) ].

Fusus (Euthria) puschi Andr. —Hoernes & Auinger 1890: 259 (pars) [non Euthria puschii ( Andrzejowski, 1830) ].

Euthria (Euthria) puschi (Andrzejowski) — Schultz 1998: 68, pl. 27, fig. 3 [non Euthria puschii ( Andrzejowski, 1830) ].

Type material. Holotype: NHMW 1872 /0005/0027a, SL: 53.9 mm, MD: 26.7 mm, Grund ( Austria), Figs 36A 1 – A 2 ; Paratypes: NHMW 1872 /0005/0027b, SL: 52.5 mm , MD: 29.8 mm, Grund ( Austria) , Figs 36B 1 –B 2 ; NHMW 1872 /0005/0027c, SL: 46.9 mm , MD: 23.4 mm, illustrated in Schultz (1998: pl. 27, fig. 3), Grund ( Austria) , Figs 36C 1 –C 2 ; NHMW 1872 /0005/0027d, SL: 47.7 mm , MD: 25.5 mm, Grund ( Austria) , Figs 36D 1 –D 2 ; NHMW 1851 /0002/0044, Grund ( Austria) . NHMW 2023 /0342/0001, SL: 48.1 mm , MD: 25.2 mm, Grund ( Austria) , Figs 36E 1 –E 2 .

Additional material. 2 spec., NHMW 1872 /0005/0027e, Grund ( Austria) ; 8 spec., NHMW 1860 /0001/0228, Mikulov ( Czech Republic) .

Type locality. Grund ( Austria), North Alpine-Carpathian Foreland Basin .

Type stratum. Silty clay of the Grund Formation.

Age. Middle Miocene, early Badenian (Langhian).

Etymology. In honor of Giano Della Bella ( Italy), in recognition for his contributions on Mediterranean Neogene malacology.

Diagnosis. Medium-sized, inflated ovate shell with conical spire, weakly incised suture and faint shoulder with low, widely spaced tubercles on last three whorls, columella and outer lip smooth, siphonal canal moderately short, recurved, deeply notched.

Description. Medium-sized, inflated ovate shell with conical spire, consisting of up to seven teleoconch whorls; apical angle 60–65°. Protoconch poorly preserved, high conical of about two convex whorls. Early teleoconch whorls with very broad, weakly concave subsutural ramp, weakly angled shoulder placed short distance above suture, most of whorl below covered by subsequent whorl. Sculpture of low, inconspicuous, widely spaced, axial ribs, overrun by indistinct spiral cords, two more prominent spirals placed at periphery and close above abapical suture. On second and third teleoconch whorls axials weaken and become wider spaced, fading thereafter. Weak primary spiral cords and indistinct secondaries on early teleoconch whorls fading on last two spire whorls. Later spire whorls with very broad, flat to faintly concave subsutural ramp and weak shoulder placed below mid-whorl bearing low, widely spaced, tubercles. Suture weakly incised. Last whorl high, attaining ~75% of total height, ovate, smooth except for 12–13 low, widely spaced tubercles placed along weakly angled shoulder; base moderately constricted with weak, widely spaced spiral cords; fasciole indistinct. Aperture wide, ovate. Columella broadly excavated, weakly angled at transition to siphonal canal. Columellar callus forming indistinct rim, poorly delimited from base, smooth. Anal canal weakly incised, without parietal and anal denticles. Outer lip not thickened, smooth within. Siphonal canal moderately short, wide, deflected to the left, strongly recurved, deeply notched.

Discussion. This species was confused with Euthria puschii ( Andrzejowski, 1830) by Hörnes (1853), Hoernes & Auinger (1890) and Schultz (1998). They differ in the more inflated shape and weaker spiral sculpture of Euthria dellabellai nov. sp. and the presence of lirae inside the outer lip in E. puschii . In addition, E. puschii has stronger tubercles at the shoulder and stronger spiral sculpture. Euthria gallica Peyrot, 1928 , from the Early Miocene of the Aquitaine Basin, differs in its much slenderer outline, higher spire and prominent spiral sculpture on the last whorl and base ( Peyrot 1928: pl. 5, figs 1–2).

Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al. 2005; Roetzel 2009). Preservation the fossil shells, suggest, that Euthria dellabellai belonged to the allochthonous shallow water fauna and not to the autochthonous deep-water fauna.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine-Carpathian Foreland Basin: Grund ( Austria) (hoc opus). Vienna Basin: Mikulov ( Czech Republic) (hoc opus).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 36. Euthria dellabellai nov. sp., A1–A2. NHMW 1872/0005/0027a, holotype, Grund (Austria). B1–B2. NHMW 1872/0005/0027b, paratype, Grund (Austria). C1–C2. NHMW 1872/0005/0027c, paratype, Grund (Austria). D1–D2. NHMW 1872/0005/0027d, paratype, Grund (Austria).