Euthria depressospira Bandat, 1943

Harzhauser, Mathias & Landau, Bernard M., 2024, The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea, Zootaxa 5427 (1), pp. 1-110 : 70-71

Euthria depressospira Bandat, 1943

Figs 34C , 37A–D

Fusus (Euthria) intermedius Micht. — Macovei 1909: 148, pl. 11, fig. 7 [non Euthria intermedia ( Michelotti, 1847) ].

* Euthria cornea Linné n. var. depressospira— Bandat 1943: 296, pl. 1, fig. 12.

Euthria oppenheimii n. sp. — Bandat 1943: 297, pl. 1, figs 13–14.

Euthria subnodosa Hoernes et Auinger — Csepreghy-Meznerics 1972: 28, pl. 11, figs 7–8 [non Euthria subnodosa Hoernes & Auinger, 1890 ].

Euthria intermedia (Michelotti, 1839) — Bałuk 1995: 243, pl. 34, fig. 10 [non Euthria intermedia ( Michelotti, 1847) ].

Euthria (Euthria) intermedia (Michelotti, 1839) — Mikuž 2009: 24, pl. 7, fig. 88 [non Euthria intermedia ( Michelotti, 1847) ].

Euthria sp. — Caze et al. 2010: 32, fig. 5J.

Euthria intermedia (Michelotti) — Kovács 2018: 182, figs 11–12 [non Euthria intermedia ( Michelotti, 1847) ].

Euthria curvirostris ( Grateloup, 1845) — Kovács 2022: 73, figs 26–27 [non Euthria curvirostris ( Grateloup, 1845) ].

Type material. Lectotype (designated herein): specimen illustrated in Bandat (1943: pl. 1, fig. 12), Middle or Late Miocene , Gurëz ( Albania). We are not aware of the whereabouts of the material.

Illustrated material. NHMW 1862/0001/0229, SL: 42.8 mm, MD: 21.9 mm, Baden ( Austria), Fig 37A 1 – A 2 . NHMW 1860/0040/0160, SL: 43.4 mm, MD: 22.0 mm, Lăpugiu de Sus ( Romania), Fig 37B 1 –B 2 . NHMW 2023/0343/0001, SL: 46.2 mm, MD: 22.8 mm, Baden ( Austria), Figs 37C 1 –C 2 . NHMW 2023/0338/0008, SL: 16.9 mm, MD: 8.5 mm, Lăpugiu de Sus ( Romania), Figs 34C .

Additional material. 10 spec., NHMW 1854 /0035/0211, Lăpugiu de Sus ( Romania) ; 28 spec., NHMW 1863 /0015/0180, Lăpugiu de Sus ( Romania) ; 15 spec., NHMW 1869 /0001/0493, Forchtenau ( Austria) ; 2 spec., NHMW 1872 /0030/0050, Baden-Sooss ( Austria) ; 3 spec., NHMW 1856 /0009/0011, Bad Vöslau ( Austria) ; 3 spec., NHMW 1862 /0001/0259, Möllersdorf ( Austria) ; 1 spec., NHMW 1853 /0038/0025, Korytnica ( Poland) .

Revised description. Medium-sized, solid, stout ovate shell with conical spire, of up to seven teleoconch whorls; apical angle 57–60°. Protoconch mammillate, of 1.5 strongly convex whorls; diameter: 880 μm, height: 980 μm. Early teleoconch whorls almost straight sided with periphery just above abapical suture. Sculpture of widely spaced, slightly drop-shaped axial ribs, overrun by four weak spiral cords and indistinct secondary cords. Suture weakly undulating around axial ribs. Axial ribs fading on third teleoconch whorl. Spiral sculpture reduced to narrow, widely spaced cords on third to fifth whorl, subobsolete on later whorls. Later spire whorls flat-sided or weakly convex, smooth, separated by narrow, weakly incised suture. Last whorl ovate, attaining ~75% of total height, with poorly delimited, weakly convex subsutural ramp, broadly convex mid-whorl, base weakly constricted, fasciole broad, smooth, indistinct. Aperture moderately wide, ovate. Columella strongly and broadly excavated, adherent, smooth except for weak denticle at angled transition to siphonal canal. Anal canal accentuated by low, broad parietal swelling, narrowly incised on broad adapical tip of aperture. Anal denticle usually absent; rarely represented by weak swelling. Outer lip convex, not thickened, bearing about 14 short, prominent lirae, not extending deeply within aperture. Siphonal canal moderately short, wide, strongly deflected and recurved, deeply notched.

Synonyms. Bandat (1943) introduced two names for this species. Based on specimens from the Miocene of Albania from the Proto-Mediterranean Sea. Euthria cornea var. depressospira Bandat 1943 was based on a fully-grown specimen, whereas Euthria oppenheimii Bandat 1943 represents a subadult specimen. As first revisers we chose Euthria depressospira as name for this species.

Discussion. This species is characterized by its solid, smooth, ovate shell with conical spire and indistinct suture. It was confused in the literature and in the collection of the NHMW with Euthria intermedia ( Michelotti, 1847) , Euthria subnodosa Hoernes & Auinger, 1890 and

Euthria curvirostris ( Grateloup, 1845) . Euthria intermedia , from the Burdigalian of the Colli Torinesi ( Italy), but differs in its slenderer shell, higher spire and prominent spiral cords on the base.

The Paratethyan Euthria subnodosa is smaller, has weak axial swellings and spiral cords on the last whorl. Euthria curvirostris , from the Serravallian of the Aquitanian Basin ( France), has a higher spire, a longer, strongly twisted siphonal canal and prominent spiral sculpture. Euthria viciani Kovács, 2018 , from the Badenian of Hungary, is superficially similar but differs in its cyrtoconoid spire, slightly concave subsutural ramp and longer siphonal canal. It is also similar in its stout profile and strongly reduced sculpture to Euthria fuscocingulata (Hoernes et Auinger, 1890) , but that species differs in having a longer siphonal canal, bearing slightly stronger, albeit weak, spiral sculpture, and fewer lirae within the outer lip.

Euthria obesa ( Michelotti, 1847) , from the Burdigalian of Termô-Fôurà ( Italy), might be closely related but differs in its stout, cyrtoconoid spire (see Bellardi 1972: pl. 13, fig. 13; Brunetti & Della Bella 2016: fig. 6E).

Paleoenvironment. The occurrence at Baden, Baden-Sooss and Möllersdorf suggests middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1995); Vienna Basin: Baden, Baden-Sooss, Möllersdorf ( Austria) (hoc opus); Pannonian Basin: Bánd ( Hungary) ( Kovács 2018); Bükk Mountains: Bóta ( Hungary) ( Csepreghy-Meznerics 1972); Krško Basin: Gorenje Vrhpolje ( Slovenia) ( Mikuž 2009); Bahna Basin ( Romania) ( Macovei 1909); Făget Basin: Lăpugiu de Sus, CoŞteiu de Sus ( Romania) ( Kovács 2022).

Proto-Mediterranean Sea. Middle or Late Miocene: Gurëz ( Albania) ( Bandat 1943).

Figures

FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width.

FIGURE 2. Colubraria subobscura (Hoernes & Auinger, 1884). A1–A3. NHMW 1854/0035/0170, paralectotype, Lăpugiu de Sus (Romania). B1–B2. NHMW 1859/0037/0018a, paralectotype, Lăpugiu de Sus (Romania). C1–C3. NHMW 1865/0001/0183, paralectotype, Lăpugiu de Sus (Romania). D1–D2. NHMW 1859/0037/0018b, paralectotype, Lăpugiu de Sus (Romania). E1–E3. NHMW 1874/0025/0021, lectotype, Lăpugiu de Sus (Romania). F1–F2. NHMW 2023/0338/0001, paralectotype, Lăpugiu de Sus (Romania).

FIGURE 34.A. Euthria brunettii nov.sp., NHMW2023/0338/0007,paratype, Lăpugiu de Sus(Romania).B.Euthria walaszczyki nov. sp., NHMW 1868/0001/0223, Grund (Austria). C. Euthria depressospira Bandat, 1943, NHMW 2023/0338/0008, Lăpugiu de Sus (Romania). D. Euthria diluvii (Eichwald, 1830) NHMW 1860/0001/0224b, Boršov (= Porstendorf) (Czech Republic). E. Euthria fuscocingulata (Hoernes & Auinger, 1890). NHMW 2023/0338/0011, paralectotype, Lăpugiu de Sus (Romania). F. Euthria frausseni nov. sp., NHMW 2023/0338/0012, paratype, Lăpugiu de Sus (Romania). G. Euthria odiosa nov. sp. NHMW 2023/0348/0001, paratype, Steinebrunn (Austria). H. Euthria puschii (Andrzejowski, 1830), NHMW 2023/0350/0002, Bad Vöslau (Austria). I. Euthria stuetzii (Naumann, 1852), NHMW 1846/0037/0254, Enzesfeld or Gainfarn (Austria). J. Euthria subnodosa (Hoernes & Auinger, 1890), NHMW 1846/0037/0274, Steinebrunn and Gainfarn (Austria). K. Euthria yesimae nov. sp., RGM 794 572, paratype, Pınarlar Yaylası, Akpınar (Turkey).

FIGURE 37. Euthria depressospira Bandat, 1943. A1–A2. NHMW 1862/0001/0229, Baden (Austria). B1–B2. NHMW 1860/0040/0160, Lăpugiu de Sus (Romania). C1–C2. NHMW 2023/0343/0001, Baden (Austria).