Rhytidaspis variata variata, Ingrisch, 2019

Ingrisch, Sigfrid, 2019, Revision of the genus Rhytidaspis Redtenbacher, 1891 including the description of a new genus Haudrhytidaspis gen. nov. (Orthoptera: Tettigoniidae: Conocephalinae), Zootaxa 4661 (2), pp. 343-370 : 353-354

Rhytidaspis variata variata ssp. n.

Figs. 1 D–E, 2G, 3I–J, 4E, 5K, 7N, 8K, 9E

Holotype as under R. variata sp. n.

Diagnosis. R. v. variata ssp. n. differs from both other subspecies by the large triangular and strongly projecting distal angle of the dorsal apical lobe of the male cercus that is separated by a weak but distinct constriction of the margin between the projecting proximal and distal angles, while in both other subspecies there is a short angular excision behind the projecting proximal angle. The stridulatory file on the underside of the male tegmen is longer than in both other subspecies, but the number of teeth in middle of the file is not higher than in R. v. brevis ssp. n., the subspecies with the shortest stridulatory file within R. variata .

Description. Male. Tegmen behind stridulatory area with converging lateral margins, apex rounded ( Figs. 3 I– J). Stridulatory file on underside of left tegmen 2.8 mm long, with about 97 teeth, in middle of file with 28.0 teeth per 1 mm, towards end teeth becoming very small and dense ( Fig. 4E ). Mirror on right tegmen roughly rhombic, internal margin strongly oblique, 1.63 mm wide, 1.03 mm long ( Fig. 3J ).

Cerci with dorsal apical lobe running oblique and parallel along inner margin of cercus trunk to nearly basal third of cercus, the basal most area of dorsal apical lobe is separated by a wide incision from the remainder of the lobe and narrow, finger-shaped with bifid tip while the apical area of the lobe is largely projecting behind the cercus stem and has a convex external and a little concave internal margin ( Fig. 5K ). The ventral apical lobe has the hind margin little concave and provided with an acute curved tooth at the underside of the inner half of the apical margin ( Fig. 7N ).

Titillators with granular extension above the U-shaped curvature with margin in middle sub-straight to slightly concave, at proximal end rounded, and distally prolonged into a short, in lateral view triangular projection; separated by a long arm from little curved tip of titillator, which is only in apical area covered by spinules and a few larger teeth ( Figs. 8K , 9E ).

Coloration. Light to medium red brown. Antennal scrobae, scapus and pedicellus black, flagellum yellow. Anterior abdominal tergites light brown, only along hind margin darkened, sixth to ninth abdominal tergites dark to blackish brown but not black; tenth tergite dark brown at base, afterwards, including apical cones and cerci yellow ( Figs. 1 D–E). Tegmen bright yellow with black dots, stridulatory area semitransparent. Legs yellow; femora with dark brown transverse strokes; hind femur in basal area dark brown. Thoracic sternites and coxae of all legs dark brown or black. Male subgenital plate black.

Measurements (1 male).—Body: 46; pronotum: 13; tegmen: 8.7; hind femur: 25.5; anterior femur: 13.5 mm.

Figures

FIGURE 1. A–B, Rhytidaspis picta Redt., 1891 (A, male Fakfak; B, female Arguni); C, R. ornata sp. n. (male South N.G.); D–E, R. variata variata ssp. n. male (Nabire); F–G, R. nigropunctata sp. n., male (Mt. Nok); H–I, R. camela sp. n., male (Sorong); J–K, R. arfak sp. n., (J, female Siwi; K, male Koffo); L, R. genyem sp. n. female (Genyem); M, R. variata brevis ssp. n. male (Freeport); N, Haudrhytidaspis fusca (Karny, 1911) comb. nov. female (type).—Habitus dorsal view (A, C, E–F, H, K, M–N) and lateral view (B, D, G, I–J, L). Scales 10 mm.

FIGURE 3. Left and right male tegmina of A–D, Rhytidaspis picta Redt., 1891 (A–B, Fakfak; C–D, Banda); E–F, R. arfak sp. n. (Koffo); G–H, R. variata meja ssp. n. (Tafelberg); I–J, R. variata variata ssp. n. (Nabire); K–L, R. variata brevis ssp. n. (Freeport); M–N, R. camela sp. n. (Sorong); O–P, R. nigropunctata sp. n. (Mt. Nok); Q–R, R. arfak sp. n. (Siwi); S–X, R. ornata sp. n. (S–T, South N.G.; U–V, Mindiptana; W–X, Tiel); Y, stridulatory file on underside of left tegmen of R. picta Redt., 1891 (Fakfak). Scales 5 mm, in Y 1 mm.

FIGURE 4. Stridulatory file on underside of left tegmen of A, Rhytidaspis arfak sp. n. (Koffo); B–C, R. camela sp. n. (B, Sorong; C, Komara); D, R. variata meja ssp. n. (Tafelberg); E, R. variata variata ssp. n. (Nabire); F, R. variata brevis ssp. n. (Freeport); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, South N.G.; I, Mindiptana; J, Tiel).

FIGURE 5. Male abdominal apex with cerci in dorsal view: A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. nigropunctata sp. n. (Mt. Nok); D–E, R. camela sp. n. (D, Sorong; E, Komara); F–G, R. variata meja ssp. n. (F, Dorey; G, Tafelberg); H–I, R. ornata sp. n. (H, Tiel; I, Mindiptana); J, R. variata brevis ssp. n. (Freeport); K, R. variata variata ssp. n. (Nabire).

FIGURE 7. Male abdominal apex or only cercus in lateral and ventral view: A–C, Rhytidaspis arfak sp. n. (A–B, Siwi; C, Koffo); D–E, R. picta Redt., 1891 (Fakfak); F–G, R. ornata sp. n. (South N.G.); H–J, R. camela sp. n. (H, Komara; I–J, Sorong); K–M, R. ornata sp. n. (K–L, South N.G.; M, Tiel); N, R. variata variata ssp. n. (Nabire); O–Q, R. variata meja ssp. n. (O–P, Tafelberg; Q, Dorey); R–S, R. variata brevis ssp. n. (Freeport); T–U, R. nigropunctata sp. n. (Mt. Nok).—Ventral view (A, E–I, K, P, S, U), oblique lateral view (B–D, J, L–O, Q–R, T).

FIGURE 8. Male titillators in lateral or oblique lateral view (A in internal, B in external view; E left in internal, right in top view): A–C, Rhytidaspis arfak sp. n. (A–B, Siwi; C, Koffo); D, R. picta Redt., 1891 (Fakfak); E–F, R. camela sp. n. (E, Sorong; F, Komara); G, R. nigropunctata sp. n. (Mt. Nok); H–J, R. ornata sp. n. (H, Tiel; I, Mindiptana; J, South N.G.); K, R. variata variata ssp. n. (Nabire); L–M, R. variata meja ssp. n. (L, Tafelberg; M, Dorey); N, R. variata brevis ssp. n. (Freeport).—Abbreviations: ap apical projection of granular sac, as oval apico-lateral sclerite, at apical branch of titillator, ea elongate apical sclerite, gs granular sac on top of U-shaped fold, ms basal muscular sac, ti base of titillator.—Tip of titillator is on top (A–F, H–K); tip is right (G, L–N).

FIGURE 9. Male titillators in ventro-apical view (tip on top): A, Rhytidaspis picta Redt., 1891 (Fakfak); B, R. arfak sp. n. (Koffo); C, R. variata meja ssp. n. (Tafelberg); D, R. variata brevis ssp. n. (Freeport); E, R. variata variata ssp. n. (Nabire); F, R. camela sp. n. (Sorong); G, R. nigropunctata sp. n. (Mt. Nok); H–I, R. ornata sp. n. (H, Mindiptana; I, South N.G.).—Abbreviation: fc flexible connection between both titillators.