Ramuliseta dolini V. Korneyev, 2015

Korneyev, V. A., 2015, A New Species Of The Genus Ramuliseta (Diptera, Ctenostylidae) From Madagascar, With A Key To Species, Vestnik Zoologii 49 (6), pp. 489-496 : 490-492

Ramuliseta dolini V. Korneyev , sp. n. ( fig 1, 1 ; 2 )

Material. Type. Holotype ♀: Madagascar: Andasibe , Perinet Nat. Reserve , 18°50ʹ S 48°23ʹ E, h = 1200 m, at light, 30.12.2003 – 8.01.2004 (Dolin & Andreeva) ( SIZK).

Diagnosis. Moderately large fly with yellow body and its vestiture, and wing with yellow base, two hyaline incisions in the costal cell, two triangular hyaline incisions at anterior margin distal to vein R 1 reaching R 2+3 and R 4+5 vein, correspondingly, and 3 large hyaline incision at posterior margin, separated by yellow bands: two from cell cua1 (basal reaching CuA 1 vein and distal entering through cell dm into r 4+5) and the apical incision covering almost all cell m and widely entering into cell r 4+5; wing apex is covered by moderately wide yellow crossband. It is similar to R. madagascariensis Hennig 1960 , in most features, differing by the wing pattern as noted in the key.

Description. Head. Length: height: width ratio = 1: 1: 1.1. Unicolorous yellow; microtrichia often indistinguishable in the holotype. Frons densely yellow setulose, except bare postero-medial ¼ of frons length bare; anteriormost setulae proclinate, middle inclinate, and posteriormost ones reclinate. Vertical plate sparsely microtrichose, with 1–2 orbital setae (1 pro- and 1 re-lateroclinate setae at left and 1 re-lateroclinate). Ocellar triangle indistinguishable; only 1 unpaired ocellar seta as long as orbital setae present. Medial vertical setae well-developed, lateral vertical setae lacking. Face membraneous, flat, matt-grey, non-translucent, with low medial carina in dorsal half and epistome slightly produced anteriorly. Parafacial and gena matt, smooth, yellow setulose. Antenna yellow, scape long and densely yellow setulose dorsally; pedicel without notch or incision, long and densely yellow setulose on its medial surface and somewhat shorter and sparser setulose on lateral surface. Flagellomere 1 yellowish, rather short, wide at base, narrow, finger-like in apical 0.7 of its length, with arista inserted into dorsal surface at its base. Arista strongly branching at base into 2–3 main stems, with numerous further dendritical branchings. Mouthparts rudimentary. Palp present, narrow and long, ventrally setulose.

Thorax. Ground colour yellow. Thorax 1.1 times as long as wide. Scutum measured between notopleural setae almost as long as wide, bearing: 0 acrostichal, 2–3 dorsocentral, 2 posterior notopleural, 1 intraalar and 1 postalar seta. Postpronotal lobe without setae, but 1–2 setae and 4–5 smaller setulae. Notopleuron without setulae. Scutellum 2.5 times as wide as long, with 2 pairs of setae (basal and apical) and with at least 2 pairs of marginal setulae between them and 6 irregular setulae on disk. Prosternum transverse, sclerotised at posteroventral margin. Proepisternum without setae or setulae. Anepisternum wrinkled in antero-ventral portion, dorsally setulose, at posterior margin with 2 setae and 7–8 setulae. Katepisternum with strong setulae inserted in antero-ventral area, medially of fore coxa margin; postero-ventral margin adjacent to mid coxa with 4–5 long setulae; latero-dorsal area with 4–5 subequal setae and setulae. Anepimeron with 6–7 subequal setae and setulae.

Legs yellow, with yellow setulae. Fore coxa robust, slightly compressed in anterodorsal direction, with moderately long yellow setae and setulae. Fore trochanter with two longer dorsal setulae. Fore femur and tibia without strong and long setae. Fore tarsus as long as tibia, yellowish-brown setulose; tarsomere 2 0.65 times as long as tarsomere 1. Mid coxa conspicuously smaller than fore and hind coxa. Mid trochanter with 2 setulae on dorsolateral surface. Mid femur, tibia and tarsus yellow setulose 6; tibia without any thickened setae at apex, with two constrictions. Hind coxa large, almost cylindrical, finely setulose on anterior surface and on sides, posterior surface mostly bare. Hind trochanter finely setuloseon posterior surface. Hind femur without setae, tibia with two constrictions.

Wing as on fig. 2 , 6. Costa thickened, with costagial break, indistinct humeral break and broad subcostal break. Humeral vein joining subcostal vein somewhat distad of its separation from R 1. Subcostal vein broken at apex, not reaching costa. Vein R 1 setulose along its whole length on dorsal side; cell r 2+3 apically widened; vein R 4+5 bare; vein M almost parallel to R 4+5; ratio of its three last sections: M2: M3: M4 = 1: 0.8: 1.3, i. e., penultimate section shorter than ultimate and second one. Wing pattern, as in most ctenostylines, with yellow basal cells and well-developed chelate pattern of connected yellow crossbands separated by hyaline incisions in apical two thirds: two hyaline incisions in the costal cell, two triangular hyaline incisions at anterior margin distal to vein R 1 reaching R 2+3 and R 4+5 vein, correspondingly, and 3 large hyaline incision at posterior margin, separated by yellow bands: two from cell cua1 (basal reaching CuA 1 vein and distal entering through cell dm into r 4+5) and the apical incision covering almost all cell m and widely entering into cell r 4+5; wing apex is covered by moderately wide yellow crossband. Vein CuA 2 +A 1 straight, in apical 1/6 of its length fold-like, but visible to its extreme tip. Vein A 2 long. Alula moderately wide, entirely hyaline. Upper calypter narrow, with clutch of long yellow setulae ventrally; lower calypter narrow. Haltere yellow, with large sparsely microtrichose knob; L-shaped.

Abdomen yellow. Syntergite 1+2 yellow, 1.7 times as long as wide, slightly longer than tergites 3–6 combined; tergites 5 and 6 with large laterobasal black spots. Sternites yellow, black setulose. Sternite 1 longitudinal trapezoid, as well as sternite 2, with clear seam between them; sternites 3–6 subquadrate, poorly sclerotized. Membrane without any setulae, short microtrichose. Oviscape convex, yellow, yellow setulose, as long as tergites 3–6 combined. Oviscape simple, tubular, with microtrichose cercal unit, flexibly joined to its apico-dorsal surface

Measurements. Body 9.5 mm, wing 8.7 mm.

Male unknown.

Etymology. This species is named in memory of its collector, famous Ukrainian entomologist Prof. Vladimir G. Dolin (1932–2004).

Figures

Fig. 1. Ramuliseta, wing: 1 — R. dolini sp.n.; 2 — R. madagascariensis; 3 — R. palpifera (arrows show key characters).

Fig. 2. Ramuliseta dolini sp. n., holotype ♀: 1 — habitus, lateral view; 2–5 — head (2 — left, 3 — anterior, 4 — dorsal view); 6 — wing; 7 — mesonotum, dorsal view.