Phasia campbelli ( Miller, 1923 ) Sun & Marshall, 2003

Sun, Xuekui & Marshall, Stephen A., 2003, Systematics of Phasia Latreille (Diptera: Tachinidae), Zootaxa 276 (1), pp. 1-320 : 28-30

3.2.3 Phasia campbelli ( Miller, 1923) , comb. nov.

(Figures I­13, II­1.2)

Campbellia campbelli Miller, 1923: 433 ­ Townsend 1938: 43 ­ Dugdale 1969: 618, 620, 635 (diagnosis and illustrations) ­ Cantrell and Crosskey 1989: 735 (catalog).

Campbellia cockaynei Miller, 1923: 436 , syn. nov.

Hyalomyia lancifera Malloch, 1930: 309 , syn. nov.

Campbellia lancifera: Cantrell and Crosskey 1989: 735 (catalog).

Campbellia cockaynei: Cantrell and Crosskey 1989: 735 (catalog).

DESCRIPTION

Body length: 3­8 mm.

MALE: HEAD. Head compressed anteriorly. Eyes separated by a distance narrower than ocellar triangle, but wider than ocellus. Fronto­orbital plate black with grey pruinosity. Frontal vitta black, triangular. Frontal vitta at base of antennae 1.6­1.7 times as wide as fronto­orbital plate anteriorly. Ocellar setae present but fine; outer vertical setae present; inner vertical setae absent. Face black with grey pruinosity. Parafacial black with grey pruinosity, bare, and as wide as first flagellomere. Lower margin of face orange yellow, grey pruinose. Vibrissa well differentiated; intervibrissal distance 1.5­1.6 times distance between vibrissa and eye on same side; facial ridge with bristles on lower 1/5. Gena black with grey pruinosity; hairs white; height 0.1­0.12 times eye height. Lunule black and shining. Antenna black; first flagellomere 1.2 times as long as pedicel; arista thickened on basal 0.66. Length of oral opening 3­3.5 times its width. Occiput flattened, greyish yellow pruinose; hairs white. Palpus yellow or brown.

THORAX. Mesoscutum with a long and broad central longitudinal pruinose vitta, and two lateral shining black vittae; postsutural area yellow pruinose, presutural area grey pruinose, with fine black hairs. 0+0(1) acrostichal seta; 0+1(2) dorsocentral setae; 1­2 postpronotal setae; presutural supra­alar seta absent, or present, but fine; 0 postsutural intraalar seta; 1­2 notopleural setae; 1 supra­alar seta; 2 postalar setae. Pleuron thinly grey pruinose; hairs black, and with scale­like setae. Anepimeral setae medium size, black; 2 katepisternal setae; 8 meral setae. Scutellum black, shining, without pruinosity, with one pair of marginal setae; apical setae absent; discal setae arranged in one row. Subscutellum very prominent. WING. Lower calypter brown to black. Wing base with scale­like setae. Tegula black. Basicosta black. Wing broad, pictured on basal 1/2 only; petiole of apical cell 0.25­0.3 times as long as preceding section of R 4+5; M meeting R 4+5 almost at right angle. Halter brown. LEGS. Fore femur brown to black, with white or yellow hairs. Fore tibia black; without bristles, apically with 1 pv, 1 p, and 1 d. Fore tarus black. Fore claws brown, apex black; 1.1 times as long as fifth tarsomere. Pulvilli brown. Mid femur brown to black with yellow base; ventrally and laterally with whitish yellow hairs, dorsally with black hairs. Mid tibia black; with 1 ad and 1 v, apically with 1 a d, 1 a, 1 av, 1 pv, 1 p and 1 d. Hind femur brown to black with yellow base; ventrally and laterally with whitish yellow hairs, dorsally with black hairs; scale­like setae present. Hind tibia black, with 4­5 ad, 3­4 pd and 1 a; first and second hind tarus with dense fine yellow hairs.

ABDOMEN. Abdominal tergites black, longitudinal vitta absent; tergites with large purple area, extending to tergite V. Relative length of abdominal tergites I+II:III:IV:V = 1.2:1:1:1.3; relative width I+II:III:IV:V = 0.3:1:1.1:1. Syntergite I+II shiny, not pruinose. Tergite III shiny. Pruinosity of tergite IV silvery. Tergite V with V­shaped pruinose posterolateral area, extending to tergite IV; hair spots distinct. TERMINALIA. Syncercus with a shallow notch posteriorly; apex bent downward. Surstylus slender, straight, longer than cercus. Ejaculatory apodeme long, stick­like. Hypandrium broad, shorter than phallapodeme, but longer than phallus. Pregonite reduced, even apically, haired ventrally. Postgonite long, pointed, with fine hairs ventrally. Phallus broad, with fine hairs dorsally on basal 1/2. Distiphallus membranous, with three small sclerotized teeth.

FEMALE. Usually smaller than male. Wing hyaline. Eyes separated by a distance less than width of single ocellus; almost touching. Indistinct pruinosity. Anterior spine­like setae of hind tibia present. Abdominal tergites black, shining. Sternite VII (sheath) slightly longer than sternite VI; posterolaterally flattened, knife­like; broad apically; slightly bent, apex directed ventrally; sternite VII with linear wrinkles ventrally. Ovipositor bent slightly downward.

TYPE MATERIAL

Campbellia campbelli Miller, 1923 . Holotype ♂, [ NEW ZEALAND], Blackball, J.M. Campbell (type depository unknown or lost, not examined).

Campbellia cockaynei Miller, 1923 . Holotype ♂, [ NEW ZEALAND], Wellington, 1.iii.1910, L. Cockayane (BMNH, examined).

Hyalomyia lancifera Malloch, 1930 . Holotype ♂, [ NEW ZEALAND], Kennedy’s Bush, 29.i.1925, A. Tonnoir (not examined); allotype ♀, [ NEW ZEALAND], Kennedy’s Bush, 29.i.1925, A. Tonnoir (USNM, examined).

OTHER MATERIAL EXAMINED

NEW ZEALAND: DNICO Dunback, 8­18.ii.1978, B. Barratt (1♀, NZAC); Taieri, D. Miller (1♀, NZAC); MC, Ashburton, Valetta Bridge, 4.iii.1991, host: Nysius sp. , M. Stufkens (1♀, NZAC). South Island, Tasman Valley, W Lake Pukaki, 2,000ft, 2.ii.1972, W.J. Knight and P.S. Broomfield (1♀, BMNH); Rocklands, 850m, 3.iv.1979, J.S. Dugdale (1♀, NZAC); Paiaka, 12.ii.1950, and 15.xii.1950, R.A. Cumber. (7♂, NZAC, CNCI).

HOST

Nysius sp. ( Hemiptera , Lygaeidae ) ­ New Zealand (reared by M. Stufkens).